Enhypnon punctatum Carter

• Main Authors: Turco, Federica, Ślipiński, Adam, Lambkin, Christine L.
• Format: Text
• Language: unknown
• Published: Zenodo 2013
• Subjects: Biodiversity; Taxonomy; Animalia; Arthropoda; Insecta; Coleoptera; Zopheridae; Enhypnon; Enhypnon punctatum; Griffith; Weir; Murchison; Sandspit; Antarc*; Antarctica
• Online Access: https://dx.doi.org/10.5281/zenodo.6153984; https://zenodo.org/record/6153984

Enhypnon punctatum Carter (Figs 2 f, 4 f, 5 f, 6 f, 7 e) Enhypnon punctatum Carter, 1927: 231. Types. Holotype. Tasmania, Mount Wellington, H.H.D. Griffith, coll. Carter [Ga: 42.883 S 147.233 E] (AMSA, K. 67211). Other specimens examined. Tasmania: Mt. Wellington, Lea [Ga: 42.883 S 147.233 E] (1, SAMA); Sandspit River, 42.09 S 147.55 E, rainforest, Tube 149, pyrethrin knockdown Atherospermum , 2.vi. 1989, J. Diggle & P. Greenslade (1, ANIC); Scotts Peak Road, DDRFB 18 C, Pitfall trapping, FT44134, 1.iii.03, D.Driscoll, 43.0163 S 146.2254 E (1, FTas); South Picton River area, South Picton River area (Baker), Pitfall trapping, FT44088, 2.ii.08, S. Baker, 43.1797 S 146.654 E (1, FTas); Warra: Manuka Road, SSTBIG 100, Pitfall trapping, FT9567, 18.ii.00, R. Bashford, 43.07 S 146.67 E (1, FTas); same locality, SSTBIG 100, Pitfall trapping, FT28547, 17.xii. 99, R. Bashford, 43.07 S 146.67 E (1, FTas); same locality, SSTBIG 170, Pitfall trapping, FT8577, 16.ii. 99, R. Bashford, 43.07 S 146.67 E (1, FTas); same locality, SSTBIG 254, Pitfall trapping, FT9519, 24.i.00, R. Bashford, 43.07 S 146.67 E (1, FTas); same locality, SSTBIG 170, Pitfall trapping, FT9460, 17.xii. 99, R. Bashford, 43.07 S 146.67 E (1, FTas); same locality, SSTBIG 170, Pitfall trapping, FT8667, 19.iii. 99, R. Bashford, 43.07 S 146.67 E (1, FTas); same locality, SSTBIG 170, Pitfall trapping, FT8674, 19.iii. 99, R. Bashford, 43.07 S 146.67 E (1, FTas); same locality, SSTBIG 170, Pitfall trapping, FT9460, 17.xii. 99, R. Bashford, 43.07 S 146.67 E (1, FTas); same locality, SSTBIG 170, Pitfall trapping, FT8308, 17.xi. 98, R. Bashford, 43.07 S 146.67 E (1, FTas); same locality, Yee site WR, WRLET 2, Emergence trapping from log of Eucalyptus obliqua , FT29927, 9.x.01, M. Yee, 43.0935 S 146.713 E, 110, Large log, Eucalyptus obliqua (1, FTas); same locality, WRLDOG 3.3, Emergence trapping from log of Eucalyptus obliqua , FT27654, 7.v.03, R.Bashford, 43.07 S 146.67 E, Old-growth, Eucalyptus obliqua (1, FTas); same locality, WRLDOG 4.4, Emergence trapping from log of Eucalyptus obliqua , FT13954, 12.iii.02, R. Bashford, 43.07 S 146.67 E, Old-growth, Eucalyptus obliqua (1, FTas); same locality, WRLDRG 1.3, Emergence trapping from log of Eucalyptus obliqua , FT13422, 18.x.01, R. Bashford, 43.07 S 146.67 E, Regrowth, Eucalyptus obliqua (1, FTas); same locality, Yee site WR, WRLET 2, Emergence trapping from log of Eucalyptus obliqua , FT29925, 28.ii.02, M. Yee, 43.0935 S 146.713 E, 110, Large log, Eucalyptus obliqua (1, FTas); same locality, Yee site WR, WRLET 2, Emergence trapping from log of Eucalyptus obliqua , FT29920, 15.xi.00, M. Yee, 43.0935 S 146.713 E, 110, Large log, Eucalyptus obliqua (1, FTas); same locality, Yee site M, MSET 3, Emergence trapping from log of Eucalyptus obliqua , FT29298, 28.ii.02, M. Yee, 43.0933 S 146.6442 E, 110, Small log, Eucalyptus obliqua (1, FTas); same locality, Yee site M, MLET 1, Emergence trapping from log of Eucalyptus obliqua , FT29228, 28.ii.02, M. Yee, 43.0933 S 146.6442 E, 110, Large log, Eucalyptus obliqua (1, FTas); same locality, Yee site M, MLET 3, Emergence trapping from log of Eucalyptus obliqua , FT29247, 10.i.02, M. Yee, 43.0933 S 146.6442 E, 110, Large log, Eucalyptus obliqua (1, FTas); same locality, WRLDRG 5.1, Emergence trapping from log of Eucalyptus obliqua , FT31496, 17.ii.06, R. Bashford, 43.07 S 146.67 E, Regrowth, Eucalyptus obliqua (1, FTas); same locality, WRLDOG 6.4, Emergence trapping from log of Eucalyptus obliqua , FT32602, 19.ii.07, R. Bashford, 43.07 S 146.67 E, Old-growth, Eucalyptus obliqua (1, FTas); same locality, WRLDOG 1.2, Emergence trapping from log of Eucalyptus obliqua , FT30874, 25.i.05, R.Bashford, 43.07 S 146.67 E, Old-growth, Eucalyptus obliqua (1, FTas); same locality, WRLDOG 2.2, Emergence trapping from log of Eucalyptus obliqua , FT31304, 16.xi.05, R.Bashford, 43.07 S 146.67 E, Old-growth, Eucalyptus obliqua (1, FTas); same locality, WRLDRG 4.1, Emergence trapping from log of Eucalyptus obliqua , FT31419, 5.i.06, R. Bashford, 43.07 S 146.67 E, Regrowth, Eucalyptus obliqua (1, FTas); same locality, WRLDOG 4.3, Emergence trapping from log of Eucalyptus obliqua , FT31318, 16.xi.05, R. Bashford, 43.07 S 146.67 E, Old-growth, Eucalyptus obliqua (1, FTas); same locality, WRLDOG 6.4, Emergence trapping from log of Eucalyptus obliqua , FT32514, 22.i.07, R. Bashford, 43.07 S 146.67 E, Old-growth, Eucalyptus obliqua (1, FTas); same locality, WRLDOG 2.1, Emergence trapping from log of Eucalyptus obliqua , FT30852, 21.xii.04, R. Bashford, 43.07 S 146.67 E, Old-growth, Eucalyptus obliqua (1, FTas); same locality, WRLDOG 1.4, Emergence trapping from log of Eucalyptus obliqua , FT31750, 22.v.06, R. Bashford, 43.07 S 146.67 E, Old-growth, Eucalyptus obliqua (1, FTas); same locality, WR011B (Baker), Pitfall trapping, FT44082, 1.vii.01, S. Baker, 43.05 S 146.70 E (1, FTas); same locality, SSTBIG 170, Pitfall trapping, FT9579, 18.ii.00, R. Bashford, 43.07 S 146.67 E (1, FTas); same locality, SSTBIG 170, Pitfall trapping, FT9506, 24.i.00, R. Bashford, 43.07 S 146.67 E (1, FTas); Warra-Weld alt transect, A03, WR0300PMT, Pitfall trapping, FT5752, 18.xii.01, N. Doran & R. Bashford, 43.0347 S 146.6945 E, 315 m (1, FTas); same locality, A02, WR0200PLO, Pitfall trapping, FT5927, 27.ii.02, N. Doran & R. Bashford, 43.0321 S 146.7006 E, 190 m (1, FTas); same locality, A05, WR0500PLO, Pitfall trapping, FT5857, 22.i.02, N. Doran & R. Bashford, 43.0374 S 146.6883 E, 535 m (1, FTas); same locality, A05, WR0500PLO, Pitfall trapping, FT133, 30.iii.01, N. Doran & R. Bashford, 43.0374 S 146.6883 E, 535 m (1, FTas); same locality, D07, WD0700P 45 U, Pitfall trapping, FT64, 26.ii.01, N. Doran & R. Bashford, 42.9996 S 146.6154 E, 739 m (1, FTas); Warra: Warra Rd, WR011B (Baker), Pitfall trapping, FT44073, 1.ii.01, S. Baker, 43.05 S 146.70 E (1, FTas); same locality, Pitfall trapping, FT44083, 1.xi.01, S. Baker, 43.05 S 146.70 E (1, FTas); Tarraleah: Hornes Dam area, HornesPF 8.4, Pitfall trapping, FT22747, 16.xi. 92, R. Brereton, 42.274 S 146.3641 E, 740, Logged, Eucalyptus delegatensis forest (1, FTas). Diagnosis. Among the species with raised tubercles on elytra, Enhypnon punctatum is distinct in having a strongly compact and convex body and distinctly raised tubercles on elytra and pronotum Redescription. Body length: 3.45 – 3.95 mm; body maximum width (elytra): 1.7 – 2.1 mm. Body: compact, distinctly convex and comparatively stout; colour dark brown with lighter spots on elytra (Figs 2 f, 4 f); vestiture consisting of recumbent yellowish setae, very dense and thick, longer and filiform around antennomeres III – XI. Head: surface coarsely sub-rugose; maximum width at eyes (0.98 – 1.13 mm), abruptly constricted behind eyes; frons widely concave between eyes; frontal ridge relatively sinuate in frontal view, thicker in front of eyes above antennal insertion; clypeus anteriorly slightly curved, not distinctly separated from frons, relatively wide (ratio clypeus width/head max. width: 0.49 – 0.51) and with obtusely rounded lateral angles; eyes small, very strongly protruding and coarsely faceted; antenna 11 -segmented with a loose and narrow 2 -segmented club; antennomere I thick and elongate, almost twice as long as wide; II distinctly elongate (about three times as long as wide), thinner than I and longer than III; II – VII elongate and progressively shorter; VIII – X cupuliform and increasing in size; X about 1.5 times as wide as IX; XI about as wide as X and slightly pointed. Last maxillary palpomere flat, short and wide, less than 1.5 times as long as wide (see Fig. 6 f). Pronotum: widest at anterior third, only weakly explanate and gently constricted posteriorly; surface coarsely sub-rugose with large tubercles in anterior half of disc (Figs 2 f, 4 f); margins obsolete and not visible from above, barely traceable in anterior half in lateral view. Leg: protibia flattened, distinctly larger than meso- and metatibia and strongly concave on the internal side; tarsomeres not transverse. Elytra: very distinct tubercles on intervals 3 (four), 5 (four) and 7 (three), one small tubercle on interval 9; second tubercle from base very low on intervals 3 and 5 (Figs 2 f, 4 f). Distribution. Tasmania (Fig. 7 e). Enhypnon simplex Turco and Ś lipi ń ski sp. n. (Figs 3 a, 4g, 5g, 6g, 7 f) Types. Holotype. Tasmania, Hartz Mts. NP, Hartz Rd, 740 m, 8 – 10.ii. 80, Eucalyptus - Nothofagus berlesed from leaf litter, A. Newton & M. Thayer [Ga: 43.233 S 146.767 E] (ANIC). Paratypes. same data as Holotype (2, ANIC). Other specimens examined. Tasmania: Scotts Peak Road, 42.5738 S 146.223 E, 1.iii.03, DDRFB 3 A, Pitfall trapping, FT 44095, D. Driscoll (1, FTas). Diagnosis. Among the species with low tubercles on elytra, E. simplex is similar to E. laticeps in having lateral angles of clypeus acute and pronotal lateral margins obsolete. It can be distinguished by the frons weakly concave, frontal ridge only slightly sinuate and pronotum convex, without a medial depressed area. Description. Body length: 2.15 – 2.3 mm; body maximum width (elytra): 1.1 – 1.15 mm. Body: small, compact and distinctly convex; colour uniformly light brown; vestiture consiting of sparse, fine and short recumbent setae, longer around antennomeres III – XI. Head: surface coarsely sub-rugose; maximum width at eyes (0.68 – 0.70 mm), abruptly constricted behind eyes; frons weakly concave between eyes; frontal ridge slightly sinuate in frontal view, thicker in front of eyes above antennal insertion; clypeus narrow (ratio clypeus width/head max. width: 0.43 – 0.44), distinctly separated and with acute lateral angles; eyes small, protruding and coarsely faceted; antenna 11 -segmented with a loose and weakly developed 2 -segmented club; antennomere I thick and about twice as long as wide and about 1.5 times as long as II; I – III progressively thinner; III as long as II; IV – VIII shorter; VIII – X cupuliform; X about 1.5 times as wide as IX; XI weakly wider than X, rounded and only slightly pointed. Last maxillary palpomere flat and distinctly wide and securiform (see Fig. 6 g). Pronotum: widest at anterior third, very weakly explanate anteriorly and constricted posteriorly; surface uniformly coarsely sub-rugose, with no medial depressed area or tubercles; margins obsolete and not visible from above. Leg: simple; tarsomeres weakly transverse. Elytra: not costate, with very wide and distinct punctures (Figs 3 a, 4g) and two almost obsolete subapical tubercles on interval 3. Etymology. The name simplex is a Latin neuter adjective ( simplex = simple) and it refers to the simple dorsum that characterises this species compared to other congeners. Distribution. Tasmania (Fig. 7 f). Enhypnon squamosum Turco and Ś lipi ń ski sp. n. (Figs 3 b, 4 h, 5 h, 6 h, 7g) Types. Holotype. Australia, Tasmania: track off Mt. Barrow Rd, 780 m, 15 – 17.ii. 1980, Nothofagus , pyrethrin-fogging tree ferns, A. Newton & M. Thayer [Ga: 41.367 S 147.417 E] (ANIC). Paratypes. Tasmania: Same data as holotype (1, ANIC); Lottah [Ga: 41.21 S 148.017 E] (1, SAMA); 4 km SE Weldborough, 450 m, 12 – 14.ii. 80, Nothofagus , pyrethrin fogging Nothofagus cunninghami bark, A. Newton & M. Thayer [GE: 41.209 S 147.948 E] (2, ANIC); W side L. St. Clair, 750 m, 25 – 29.i. 80, Nothofagus , pyrethrin-fogging tree ferns, A. Newton & M. Thayer [GE: 42.055 S 146.140 E] (5, ANIC); Lake St. Clair, 42.06 S 146.10 E, 750 m, 25 – 27.i. 1980, Pyrethrum spray tree ferns, J. Lawrence & T. Weir (3, ANIC); Mt. Barrow Rd, 890 m, 15 – 17.ii. 1980, Nothofagus berlesed from leaf litter, A. Newton & M. Thayer [Ga: 41.367 S 147.417 E] (1, ANIC); 4,4 km SE Weldborough, 460 m, 12 – 14.ii. 1980, Nothofagus , forest, A. Newton & M. Thayer [GE: 41.210 S 147.954 E] (2, ANIC); Savage R. Pipeline Rd, 41.30 S 145.20 E, 19.iv. 1989, Tube 72, pyrethrin knockdown Nothofagus , J. Diggle (1, ANIC); same locality, 20.iv. 1989, Tube 17, pyrethrin knockdown. Nothofagus , P. Greenslade (1, ANIC); Mt. Victoria, 41.20 S 147.50 E, 900 m, 23.xi. 1989, pyrethrin knockdown tree 1, Tube 297, R. Coy (1, ANIC); same locality, 25.xi. 1989, pyrethrin knockdown tree 2, Tube 239, H. Mitchell (2, ANIC); same locality, 23.xi. 1989, pyrethrin knockdown tree 2, Tube 296, R. Coy (1, ANIC); same locality, 23.xi. 1989, pyrethrin knockdown tree 3, Tube 298, R. Coy (2, ANIC); Mt. Michael, 41.11 S 148.00E, 740 m, 26.xi. 1989, pyrethrin knockdown tree 1, Tube 187, R. Coy (1, ANIC); same locality, 27.xi. 1989, pyrethrin knockdown tree 2, Tube 182, R. Coy (1, ANIC); same locality, 28.xi. 1989, pyrethrin knockdown tree 1, Tube 176, R. Coy (1, ANIC); St. Columba Falls, Berlesate Anic 912, 16.xii. 1981, G. Bornemissza [Ga: 41.317 S 147.917 E] (1, ANIC). Notes on types. The specimen from Lottah (SAMA) bears an old cotype label and an identification label by Carter as Enhypnon horridum . Nonetheless, this specimen certainly belongs to the new species E. squamosum and it is not part of the original type series of E. horridum , which consists of three specimens collected on Cradle Mountain by Lea. Other specimens examined. Tasmania: Ben Ridge Rd., 5.2 km E Telopea Rd., 41.21 S 147.40 E, 870 m, 16.i – 1.ii. 1993, A. Newton and M. Thayer, ex window-trap Nothofagus cunninghamii rainforest (1, ANIC); Mt Michael, 41.1833 S 148.0000E, FT 38038, Knockdown fogging of canopy of Nothofagus cunninghamii , 21 /02/ 90, R.Coy, P. Lillywhite & A.Yen (6, FTas); same locality, FT 38037, Knockdown fogging of canopy of Nothofagus cunninghamii , 20 /02/ 90, R.Coy, P. Lillywhite & A.Yen (9, FTas); same locality, 41.11 S 148.00E, 740 m, 27.xi. 1989, pyrethrin knockdown tree 1 – 2, Tube (ranging from 182 to 187), R. Coy (6, ANIC); same locality, 28.xi. 1989, suction, Tube 245, R. Coy (1, ANIC); same locality, 11.vi. 1990, moss on tree trunks, Tube 320, A.T. W. (1, ANIC); Mt. Victoria, 41.20 S 147.50 E, 900 m, 25.xi. 1989, pyrethrin knockdown tree 1 – 3, Tube (ranging from 237 to 305), H. Mitchell (or R. Coy) (46, ANIC); same locality, 22 – 28.xi. 1989, yellow pans (x 5), Tube 169, H. Mitchell & R. Coy (1, ANIC); same locality, 29.xi. 1989, litter, Tube 289, R. Coy (1, ANIC); 4,4 km SE Weldborough, 460 m, 12 – 14.ii. 1980, Nothofagus , forest, A. Newton & M. Thayer [GE: 41.210 S 147.954 E] (1, ANIC); Savage R. Pipeline Rd, 41.30 S 145.20 E, 19.iv. 1989, Tube 72, pyrethrin knockdown Nothofagus , J. Diggle (1, ANIC); Schulhofs Rd., 8.7 km NE Upper Blessington, 41.26 S 147.36 E, 870 m, 16.i – 1.ii. 1993, A. Newton & M. Thayer, ex window-trap, Euc. delegatensis and Euc. obliqua forest (1, ANIC); Simons Rd., non-wilt area, 41.21 S 147.31 E, 8.iii. 1989, leaf litter, Tube 273, P. Greenslade (4, ANIC); Simons Rd., wilt area, 41.21 S 147.31 E, 8.iii. 1989, leaf litter, Tube 272, J. Diggle (1, ANIC); Near Weldborough, 41 ° 10 ’S 147 °08’E, 20 – 25.ii. 1993, Nothofagus forest, litter, J. Trueman et al. , site A (1, ANIC); 40 km SW Smithton, 4.iii. 1977, FMHD # 77 – 189, litter in Crown of Dicksonia antarctica , J. Kethley [GE: 41.047 S 144.734 E] (1, FMNH); Cradle Mtn. Nat. Pk, Waldheim For. (3500ft), 3.ii. 1977, FMHD # 77 – 109, litter of unburned Nothofagus gunnei , J. Kethley [Ga: 41.633 S 145.933 E] (1, FMNH); 2 km S. Revision Bell, 9.xii. 1986, D. Burckhardt (1, ANIC). Diagnosis. Among the species with raised tubercles on elytra, Enhypnon squamosum is distinct in having deeply denticulate pronotal margins and strongly squamiform dorsal vestiture. This species is quite variable in size and colour, from uniformly yellowish-brown to distinctly bicoloured with patterned elytra. Description. Body length: 1.9 – 3.0 mm; body maximum width (elytra): 1.0 – 1.7 mm. Body: small, compact and distinctly convex; colour from uniformly light brown to distinctly bicoloured with patterned elytra and lateral part of pronotum; vestiture consiting of dense, short and strongly squamiform setae (Figs 3 b, 4 h); long and filiform around antennomeres III – XI. Head: surface coarsely sub-rugose; maximum width at eyes (0.60 – 0.88 mm), abruptly constricted behind eyes; frons broadly concave between eyes; frontal ridge weakly sinuate in frontal view, thicker in front of eyes above antennal insertion; clypeus anteriorly curved, relatively wide (ratio clypeus width/head max. width: 0.49 – 0.58) and with obtuse lateral angles; eyes strongly protruding and coarsely faceted; antenna 11 -segmented with a loose and weakly developed 2 -segmented club; antennomere I relatively thin and elongate (about twice as long as wide); II more than twice as long as wide; II – VIII elongate and progressively shorter; IX more elongate and wider than VIII; X cupuliform and relatively narrow (less than 1.5 times as wide as IX); XI only slightly wider than X, sub-oval, weakly elongate and pointed. Last maxillary palpomere flat, distinctly wide and sub-triangular (see Fig. 6 h). Pronotum: explanate and abruptly constricted posteriorly; surface sub-rugose with two admedian shallow depressions on pronotal disc; margin denticulate, usually with four/five teeth or spines. Leg: simple; tarsomeres weakly transverse. Elytra: distinctly tuberculate along intervals 3, 5, 7, and 9; two larger sub-apical tubercles on interval 3, clearly visible in lateral view (Figs 3 b, 4 h). Etymology. The name squamosum is a Latin neuter adjective, squamosum (= that bears scales) and it refers to the distinctive vestiture, consisting of dense and strongly squamiform setae. Distribution. Tasmania (Fig. 7 g). Enhypnon tuberculatum Turco and Ś lipi ń ski sp. n. (Figs 3 d, 4 j, 5 j, 6 j, 7 h) Types. Holotype. Australia, Tasmania: Hartz Mts. NP, Hartz Rd, 740 m, 8 – 10.ii. 80, Eucalyptus - Nothofagus , berlesed from leaf litter, Newton & Thayer [Ga: 43.233 S 146.767 E] (ANIC). Paratypes. Tasmania: Same data as holotype (1, ANIC); 22 mi. SE Smithton, Savage Riv, 5.iii. 1977, FMHD # 77 – 193, Nothofagus cunninghami litter at base of Nothofagus cunninghami log, J. Kethley [GE: 41.103 S 145.371 E] (1, FMNH); 35 km SW Smithton, Christmas Hills, 27.ii. 1977, FMHD # 77 – 177 N. cunninghami litter around log, J. Kethley [GE: 40.901 S 144.997 E] (1, FMNH); Corinna W Darlingtons,. iii. 57 [Ga: 41.65 S 145.067 E] (1, ANIC); Mt. Murchison, 41.50 S 145.37 E, 21.iv. 1989, rainf. Tube 129 moss on : Published as part of Turco, Federica, Ślipiński, Adam & Lambkin, Christine L., 2013, Enhypnon Carter: a taxonomic revision of an endemic Australian genus of ground-dwelling beetles (Coleoptera: Zopheridae), pp. 371-394 in Zootaxa 3681 (4) on pages 385-391, DOI: 10.11646/zootaxa.3681.4.3, http://zenodo.org/record/215723 : {"references": ["Carter, H. J. (1927) Australian Coleoptera: notes and new species. No. V. Proceedings of the Linnean Society of New South Wales, 52, 222 - 234."]}