Halechiniscus remanei subsp. remanei Schulz 1955
82. Halechiniscus remanei remanei Schulz, 1955 Halechiniscus remanei nov. spec. (Schulz 1955) Halechiniscus remanei Schulz (Renaud-Debyser 1959 b, Zio de 1964, Swedmark & Teissier 1967, Fleeger 1978) Halechiniscus remanei (Zio de 1965, Grimaldi de Zio et al. 1980 a, Accogli et al. 2009) Halechin...
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| Summary: | 82. Halechiniscus remanei remanei Schulz, 1955 Halechiniscus remanei nov. spec. (Schulz 1955) Halechiniscus remanei Schulz (Renaud-Debyser 1959 b, Zio de 1964, Swedmark & Teissier 1967, Fleeger 1978) Halechiniscus remanei (Zio de 1965, Grimaldi de Zio et al. 1980 a, Accogli et al. 2009) Halechiniscus remanei Schulz, 1955 (Rao & Ganapati 1968, McGinty 1969, Renaud-Mornant 1970 c, 1971, Renaud-Mornant 1981 d, Renaud-Mornant & Gourbault 1981, 1984, Grimaldi de Zio et al. 1987, Pollock 1989, Villora-Moreno & Grimaldi de Zio 1996, Grimaldi de Zio & D’Addabbo Gallo 2001, Chang & Rho 2002, Grimaldi de Zio et al. 2003, D’Addabbo Gallo et al. 2007) Halechiniscus remanei Schulz 1955 (D’Addabbo Gallo et al. 1978, Grimaldi de Zio et al. 1979, Grimaldi de Zio et al. 1980 b) H. remanei, Schulz 1955 (Grimaldi de Zio et al. 1983 b) Terra typica : Tyrrhenian Sea (Italy, Europe) Adriatic Sea: • 42 °07′ 18 ′′N, 15 ° 29 ′ 49 ′′E; 0 m bsl: [FAO 37] Italy, Foggia Province, Tremiti Islands, San Domino Island, Cala delle Arene, intertidal, fine-medium sand, mainly calcareous. Grimaldi de Zio et al. (1983 b) • 41 ° 57 ′ 20 ′′N, 15 °03′ 23 ′′E; 0 m bsl: [FAO 37] Italy, Campobasso Province, Campomarino. Grimaldi de Zio et al. (2003) • 41 ° 55 ′ 30 ′′N, 15 ° 89 ′ 38 ′′E [41 ° 55 ′N, 15 ° 10 ′E]; 0 m bsl: [FAO 37] Italy, Foggia Province, Marina di Chieuti, intertidal, medium sand. Grimaldi de Zio et al. (1983 b) • 41 ° 33 ′ 35 ′′N, 15 ° 53 ′ 36 ′′E; 0 m bsl: [FAO 37] Italy, Foggia Province, Frattarolo, intertidal, medium-fine sand. Grimaldi de Zio et al. (1983 b) • 41 °05′N, 17 °01′E 0-3 m bsl: [FAO 37] Italy, Bari Province, Calo Colombo, coarse sand. Accogli et al. (2009) • 41 °02′ 30 ′′N, 17 °08′00′′E; 0 m bsl: [FAO 37] Italy, Bari Province, Cozze, intertidal, medium sand. Grimaldi de Zio et al. (1983 b) • 40 ° 58 ′ 45 ′′N, 17 ° 15 ′ 47 ′′E; 0 m bsl: [FAO 37] Italy, Bari Province, Torre Incina, intertidal, on the shoreline, coarse sand (0.841 mm). Grimaldi de Zio et al. (1980 b, 1983 b) • 40 ° 57 ′ 30 ′′N, 17 ° 17 ′ 30 ′′E [40 ° 58 ′N, 17 ° 16 E]; 0 m bsl: [FAO 37] Italy, Bari Province, Cala Corvina [Cala Corvino], intertidal, medium sand, mainly calcareous. Grimaldi de Zio et al. (1983 b) • 40 ° 54 ′ 25 ′′N, 17 ° 20 ′ 40 ′′E; 0 m bsl: [FAO 37] Italy, Bari Province, Capitolo Sud, intertidal, medium coarse calcareous sand. Grimaldi de Zio et al. (1983 b) • 40 ° 53 ′ 15 ′′N, 17 ° 23 ′ 30 ′′E; 0 m bsl: [FAO 37] Italy, Brindisi Province, Torre Egnazia, intertidal, medium calcareous sand. Grimaldi de Zio et al. (1983 b) • 40 ° 51 ′ 20 ′′N, 17 ° 25 ′ 12 ′′E [40 ° 50 ′N, 17 ° 28 ′E]; 0 m bsl: [FAO 37] Italy, Brindisi Province, Torre Canne, intertidal, medium-fine sand. Grimaldi de Zio et al. (1983 b) • 40 ° 50 ′N, 17 ° 28 ′E; 0 m bsl: [FAO 37] Italy, Brindisi Province, Torre Canne, Masseria Fiume Morello, beach, tidal zone to minimum level of low tide, max 6 m from the sea to the river, salinity (<30 ‰), sand. Zio de (1965) , Grimaldi de Zio et al. (1979) • 40 ° 46 ′ 20 ′′N, 17 ° 39 ′ 30 ′′E; 0 m bsl: [FAO 37] Italy, Brindisi Province, Torre Pozzelle, intertidal, medium sand. Grimaldi de Zio et al. (1983 b) • 40 ° 22 ′ 54 ′′N, 18 ° 18 ′ 30 ′′E; 0 m bsl: [FAO 37] Italy, Lecce Province, San Cataldo, beach, intertidal, mediumfine homogeneous sand (0.149–0.595 mm). Zio de (1964) , Grimaldi de Zio et al. (1983 b) Alboran Sea: • 35 ° 11 ′N, 02° 26 ′W; 0 m bsl: [FAO 37] Morocco, Chafarinas Islands, Isable II Island [Isla Isabel II], shallow sublittoral sediment between rock blocks. Villora-Moreno & Grimaldi de Zio (1996) Atlantic Ocean: • 33 ° 20 ′N, 79 ° 10 ′W [FAO 31] USA, South Carolina State, Georgetown County, North Inlet, subtidal sand. Fleeger (1978) • 33 ° 20 ′N, 79 ° 10 ′W 0 m bsl: [FAO 31] USA, South Carolina State, Georgetown County, North Inlet, intertidal mud between Spartina alterniflora plants. Fleeger (1978) • 44 ° 40 ′N, 01° 14 ′W; 0 m bsl: [FAO 27] France, Aquitaine Region, Arcachon Bay, American beach, 20–40 cm deep in sand, sand (diameter 0.25–0.30 mm). Renaud-Debyser (1959 b) • 44 ° 40 ′N, 01° 10 ′W; 1.5 m bsl: [FAO 27] France, Aquitaine Region, Arcachon Bay, d’Eyrac beach, sand. Renaud-Debyser (1959 b) • 44 °03′N, 01° 20 ′W; 0 m bsl: [FAO 27] Undefined locality, France, Aquitaine Region, coast of Landes Department. Renaud-Debyser (1959 b) Balearic Sea: • 41 ° 14 ′ 15 ′′N, 08° 59 ′ 42 ′′E [41 ° 14 ′N, 09° 10 ′E]; 0 m bsl: [FAO 37] Italy, Sardinia, Olbia-Tempio Province, Santa Reparata. Grimaldi de Zio et al. (2003) • 41 °09′ 15 ′′N, 09° 31 ′ 48 ′′E [41 °09′N, 09°07′E]; 0 m bsl: [FAO 37] Italy, Sardinia, Olbia-Tempio Province, Monte Russu, intertidal, coarse sand. Grimaldi de Zio et al. (1983 b) • 40 ° 49 ′N, 08° 30 ′E; 0 m bsl: [FAO 37] Italy, Sardinia, Sassari Province, Gulf of Asinara, medium and medium fine sand. D’Addabbo Gallo et al. (2007) • 40 ° 20 ′N, 08° 22 ′E; 0 m bsl: [FAO 37] Italy, Sardinia, Sassari or Oristano Province, Gulf of Alghero, medium sand. D’Addabbo Gallo et al. (2007) Caribbean Sea: • 16 ° 12.5 ′N, 61 ° 29.4 ′W / 16 ° 13 ′N, 61 ° 29 ′W; 0 m bsl: [FAO 31] Guadeloupe Islands, Grande-Terre Island, Le Gosier, beach, intertidal, fine sand (mean diameter 0.225 mm). Renaud-Mornant & Gourbault (1981, 1984) Celtic Sea: • 48 °01′N, 05° 44 ′W; 130 m bsl: [FAO 27] France, Brittany Region, continental shelf, near Brest, coarse sand and shelly detritus with limestone and finer siliceous sand. Renaud-Mornant (1970 c, 1971) Coral Sea: • 19 ° 2 ′S, 147 ° 30 ′E [19 ° 15 ′S, 146 ° 49 ′E]; 0 m bsl: [FAO 71] Australia, Queensland State, Townsville, beach, intertidal coralline sand. Renaud-Mornant (1981 d) East China Sea: • 33 ° 30 ′N, 126 ° 58 ′E; 2–3 m bsl: [FAO 61] South Korea, Jeju Province, Jejudo Island, Udo Islet, sublittoral, coralline sand. Chang & Rho (2002) Indian Ocean: • 17 ° 43 ′N, 83 ° 20 ′E; 0 m bsl: [FAO 57] India, Andhra Pradesh, Bay of Bengal, Waltair coast, beach, mid-water level, medium sand. Rao & Ganapati (1968) • 20 ° 11 ′S, 57 ° 16 ′E [20 ° 11 ′S, 57 ° 46 ′E]; 0 m bsl: [FAO 51] Republic of Mauritius, Flacq Region, Belle Mar Beach, intertidal, interstitial, fine well-sorted coralline sand with Foraminifera. Grimaldi de Zio et al. (1987) Ionian Sea: • 40 ° 29 ′00′′N, 17 °03′ 25 ′′E; 15 m bsl: [FAO 37] Italy, Tarento Province, Secca dell’Armeleia, subtidal, organogenic, Amphioxus debris. Grimaldi de Zio et al. (1983 b) • 40 ° 24 ′ 30 ′′N, 17 ° 13 ′ 28 ′′E; 0 m bsl: [FAO 37] Italy, Tarento Province, Lido Bruno, intertidal, medium coarse sand with high calcareous percentage (97 %). Grimaldi de Zio et al. (1983 b) • 40 ° 23 ′07′′N, 17 ° 18 ′ 29 ′′E; 0 m bsl: [FAO 37] Italy, Tarento Province, Lido Gandoli, intertidal, heterogenous sand with high calcareous content. Grimaldi de Zio et al. (1983 b) • 40 ° 22 ′03′′N, 17 ° 22 ′ 29 ′′E [40 ° 21 ′N, 17 ° 22 ′E]; 0 m bsl: [FAO 37] Italy, Tarento Province, Gabbiano, intertidal, heterogenous sand mainly calcareous (98 %). Grimaldi de Zio et al. (1983 b) • 40 ° 22 ′ 25 ′′N, 17 ° 18 ′ 18 ′′E [40 ° 21 ′N, 17 ° 22 ′E]; 0 m bsl: [FAO 37] Italy, Tarento Province, Marina di Pulsano. Grimaldi de Zio et al. (2003) • 40 ° 21 ′N, 17 ° 23 ′E; 0 m bsl: [FAO 37] Italy, Taranto Province, Marina di Pulsano, Lido conche, beach, 2.5 m of the coast line, 15–20 cm deep in sand, medium coarse sand (max 0.841 mm). D’Addabbo Gallo et al. (1978), Grimaldi de Zio et al. (1980 b) • 40 ° 17 ′ 45 ′′N, 17 ° 33 ′ 38 ′′E; 0 m bsl: [FAO 37] Italy, Tarento Province, Campo Marino. Grimaldi de Zio et al. (2003) • 40 °09′N, 17 ° 58 ′E 0-20 m bsl: [FAO 37] Italy, Lecce Province, Nard, Corvine cave, coarse sand. Accogli et al. (2009) • 40 °08′ 38 ′′N, 17 ° 59 ′00′′E; 0 m bsl: [FAO 37] Italy, Lecce Province, Santa Caterina, intertidal, medium sand, mainly calcareous. Grimaldi de Zio et al. (1983 b) • 40 °08′N, 17 ° 59 ′E; 0 m bsl: [FAO 37] Italy, Lecce Province, Santa Caterina, beach, 1 m from the shoreline, 35 cm deep in sand, medium sand (max 0.354 mm). D’Addabbo Gallo et al. (1978), Grimaldi de Zio et al. (1980 b) • 40 °03′00′′N, 18 °00′00′′E; 0 m bsl: [FAO 37] Italy, Lecce Province, near Gallipoli, Lido Conchiglie, intertidal, medium sand, mainly calcareous (97 %). Grimaldi de Zio et al. (1983 b) • 40 °02′ 50 ′′N, 17 ° 59 ′ 30 ′′E; 0 m bsl: [FAO 37] Italy, Lecce Province, near Gallipoli, Baia Verde, intertidal, medium sand, mainly calcareous. Grimaldi de Zio et al. (1983 b) • 40 °02′N, 18 °01′E; 0 m bsl: [FAO 37] Italy, Lecce Province, near Gallipoli, Baia Verde, beach, 1 m of the coast line, 50 deep cm deep in sand, sand (0.354 mm). D’Addabbo Gallo et al. (1978) • 39 ° 51 ′05′′N, 18 ° 12 ′ 20 ′′E [39 ° 50 ′N, 18 ° 12 ′E]; 0 m bsl: [FAO 37] Italy, Lecce Province, Torre Pali, intertidal, medium-fine sand, mainly calcareous. Grimaldi de Zio et al. (1983 b) • 39 ° 50 ′ 40 ′′N, 18 ° 10 ′ 25 ′′E [39 ° 51 ′N, 18 ° 11 ′E]; 0 m bsl: [FAO 37] Italy, Lecce Province, Lido Marini, intertidal, medium coarse sand. Grimaldi de Zio et al. (1983 b) • 39 ° 50 ′ 40 ′′N, 18 ° 10 ′ 15 ′′E; 50 m bsl: [FAO 37] Italy, Lecce Province, Lido Marini, subtidal. Grimaldi de Zio et al. (1983 b) Mediterranean Sea: • 36 ° 45 ′00′′N, 02° 53 ′00′′E [36 ° 46 ′N, 02° 53 ′E]; 0 m bsl: [FAO 37] Algeria, Tipaza Province, near Staouéli. Grimaldi de Zio et al. (1980 a) • 36 ° 38 ′00′′N, 02° 40 ′00′′E; 0 m bsl: [FAO 37] Algeria, Tipaza Province, near Khemisti. Grimaldi de Zio et al. (1980 a) • 35 ° 54 ′N, 14 ° 32 ′E; 41 m bsl : [FAO 37] Republic of Malta, SE coast of Malta, Sicily Channel, 1.75 km SE of the entrance of Grand Harbour and 1.4 km NW of the coastal settlement Xghajra, coarse organogenic detritus. Grimaldi de Zio & D’Addabbo Gallo (2001) North Sea: • 59 ° 17 ′N, 11 ° 21 ′E; 0 m bsl: [FAO 27] Sweden, Västra Götalands Iän Region, Kattegat, Gunnarskär, shelly sand with shells, fragments of sea urchin tests and pieces of bryozoans. Swedmark & Teissier (1967) • 58 ° 35 ′N, 11 °04′E; 0 m bsl: [FAO 27] Sweden, Västra Götalands Iän Region, Kattegat, Väderöarna, shelly sand with shells, fragments of sea urchin tests and pieces of bryozoans. Swedmark & Teissier (1967) • 58 ° 17 ′N, 11 ° 30 ′E: [FAO 27] Undefined locality, Sweden, Västra Götaland County, Bohuslän Province, Skagerrak, Gullmarsfjord. Renaud-Mornant (1971) • 58 °06′N, 11 ° 20 ′E; 0 m bsl: [FAO 27] Sweden, Västra Götalands Iän Region, Kattegat, Bonden Island near Käringön, shelly sand with shells, fragments of sea urchin tests and pieces of bryozoans. Swedmark & Teissier (1967) Pacific Ocean: • 38 °17.0′N, 123 °2.0′W [38 ° 18 ′N, 123 °03′W]; 0 m bsl: [FAO 77] USA, California State, Sonoma County, N of the parking area at Bodega Head, 0–60 cm deep in sand, well-sorted coarse or fine sand (mean phi grains = 2.545). Pollock (1989) • 36 ° 38 ′N, 121 ° 56 ′W; 0–20 cm / 0–0.2 m bsl: [FAO 77] USA, California State, Monterey County, Pacific Grove, SW of Hopkins Marine Station, sandy beach, mid- to high tide, medium grain sand. McGinty (1969) • 35 °59.0′N, 121 °30.0′W / 35 ° 59 ′N, 121 ° 30 ′W; 0 m bsl: [FAO 77] USA, California State, Monterey County, Los Padres National Forest, Mill Creek Picnic Area, Big Sur coastline, 0–60 cm deep in sand, sand. Pollock (1989) • 33 °35.0′N, 117 °53.0′W / 33 ° 35 ′N, 117 ° 53 ′W; 0 m bsl: [FAO 77] USA, California State, Orange County, Corona del Mar, Corona del Mar State Park, Little Corona Beach, 0–60 cm deep in sand, sand with shell debris (mean phi grains = 0.655). Pollock (1989) Tyrrhenian Sea: • 40 ° 50 ′N, 14 ° 16 ′E; 0 m bsl: [FAO 37] Type Locality: Italy, Province of Naples, Naples, Gulf of Naples, coarse sand. Schulz (1955) • 40 ° 36 ′N, 14 ° 52 ′E; 0 m bsl: [FAO 37] Italy, Salerno Province, Battipaglia. Grimaldi de Zio et al. (2003) • 40 ° 13 ′N, 09° 38 ′E; 0–7 m bsl: [FAO 37] Italy, Sardinia, Nuoro Province, Gulf of Orosei, coarse and medium fine sand. D’Addabbo Gallo et al. (2007) • 38 ° 44 ′N, 16 °06′E [38 ° 43 ′N, 16 °06′E]; 0 m bsl: [FAO 37] Italy, Vibo Valentia Province, Vibo Valentia, Trainiti beach. Grimaldi de Zio et al. (2003) • 38 ° 37 ′N, 15 °03′E [38 ° 38 ′N, 15 °04′E]; 1–4 m bsl: [FAO 37] Italy, Aeolian Islands, Panarea Island, Baia del Porto, subtidal zone, siliceous sand. Grimaldi de Zio & D’Addabbo Gallo (2001), Grimaldi de Zio et al. (2003) • 38 ° 27 ′N, 14 ° 55 ′E [38 ° 27 ′N, 14 ° 56 ′E]; 1.5–5 m bsl: [FAO 37] Italy, Aeolian Islands, Lipari Island, Valle Muria, subtidal zone, siliceous sand. Grimaldi de Zio & D’Addabbo Gallo (2001), Grimaldi de Zio et al. (2003) • 38 ° 26 ′N, 14 ° 55 ′E [38 ° 26 ′N, 14 ° 57 ′E]; 1.5–5 m bsl: [FAO 37] Italy, Aeolian Islands, Lipari Island, Pietra Lunga South, subtidal zone, siliceous sand. Grimaldi de Zio & D’Addabbo Gallo (2001), Grimaldi de Zio et al. (2003) • 38 °01′ 40 ′′N, 13 ° 56 ′ 38 ′′E [38 °02′N, 14 °01′E]; 0 m bsl: [FAO 37] Italy, Sicily, Palermo Province, Cafal (La Plaia), intertidal, coarse sand, mainly siliceous (93 %). Grimaldi de Zio et al. (1983 b) • 37 ° 34 ′ 50 ′′N, 12 ° 50 ′ 38 ′′E [37 ° 35 ′N, 12 ° 50 ′E]; 0 m bsl: [FAO 37] Italy, Sicily, Trapani Province, Selinunte, intertidal, medium sand, mainly calcareous. Grimaldi de Zio et al. (1983 b) Record numbers (Sea/Ocean classification): Adriatic Sea: 14, Alboran Sea: 1, Atlantic Ocean: 5, Balearic Sea: 4, Caribbean Sea: 1, Celtic Sea: 1, Coral Sea: 1, East China Sea: 1, Indian Ocean: 2, Ionian Sea: 16, Mediterranean Sea: 3, North Sea: 4, Pacific Ocean: 4, Tyrrhenian Sea: 9; total: 66. Record numbers (FAO classification): FAO 27: 8, FAO 31: 3, FAO 37: 47, FAO 51: 1, FAO 57: 1, FAO 61: 1, FAO 71: 1, FAO 77: 4; total: 66. Remarks: This is the m : Published as part of Kaczmarek, Łukasz, Bartels, Paul J., Roszkowska, Milena & Nelson, Diane R., 2015, The Zoogeography of Marine Tardigrada, pp. 1-189 in Zootaxa 4037 (1) on pages 80-84, DOI: 10.11646/zootaxa.4037.1.1, http://zenodo.org/record/233519 : {"references": ["Schulz, E. (1955) Studien an marinen Tardigraden. 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Biology of Tardigrades. Selected Symposia and Monographs U. Z I., 1. Mucchi, Modena, Italy, pp. 93 - 101.", "Pollock, L. W. (1989) Marine interstitial Heterotardigrada from the Pacific coast of the United States, including a description of Batillipes tridentatus n. sp. Transactions of the American Microscopical Society, 108, 169 - 189. http: // dx. doi. org / 10.2307 / 3226373", "Villora-Moreno, S. & Grimaldi de Zio, S. (1996) New records of marine Tardigrada in the Mediterranean Sea. Zoological Journal of the Linnean Society, 116, 149 - 166. http: // dx. doi. org / 10.1111 / j. 1096 - 3642.1996. tb 02340. x", "D'Addabbo Gallo, M., Grimaldi de Zio, S. & Sanduli, R. (2001) Heterotardigrada of two submarine caves in S. Domino Island (Tremiti Islands) in the Mediterranean Sea with the description of two species of Stygarctidae. Zoologischer Anzeiger, 240, 361 - 369. http: // dx. doi. org / 10.1078 / 0044 - 5231 - 00043", "Chang, C. Y. & Rho, H. S. (2002) Taxonomic study of marine tardigrades from Korea II. Genus Halechiniscus (Heterotardigrada, Arthrotardigrada, Halechiniscidae). The Korean Journal of Systematic Zoology, 18, 91 - 98.", "Hansen, J. G., D'Addabbo Gallo, M. & Grimaldi De Zio, S. (2003) A comparison of morphological characters within the genus Rhomboarctus (Tardigrada: Heterotardigrada) with the description of two new species. Zoologischer Anzeiger, (242) 1, 83 - 96. http: // dx. doi. org / 10.1078 / 0044 - 5231 - 00089", "D'Addabbo Gallo, M., Morone de Lucia, M. R., Grimaldi de Zio, S. & Grimaldi, P. (1978) Nuovi dati sui tardigradi del mesopsammon delle coste pugliesi. Thalassia Salentia, 8, 63 - 66.", "Renaud-Mornant, J. (1984 b) Nouveaux Arthrotardigrades des Antilles. Bulletin du Museum National d'Histoire Naturelle, Paris, Serie 4 e, 6, 975 - 988.", "Faurby, S., Jorgensen, A., Kristensen, R. M. & Funch, P. (2011) Phylogeography of North Atlantic intertidal tardigrades: refugia, cryptic speciation and the history of the Mid-Atlantic Islands. Journal of Biogeography, 38, 1613 - 1624. http: // dx. doi. org / 10.1111 / j. 1365 - 2699.2011.02533. x"]} |
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