Diplura macrura C.L. Koch 1841

• Main Authors: Pedroso, Denis Rafael, De Souza Castanheira, Pedro, Renner Luiz Cerqueira Baptista
• Format: Text
• Language: unknown
• Published: Zenodo 2016
• Subjects: Biodiversity; Taxonomy; Animalia; Arthropoda; Arachnida; Araneae; Dipluridae; Diplura; Diplura macrura; Galapagos; Greenland; Argentina; Fukushima; Seta; Bryant; Duarte; Juv; Blackwall; Blunt Point
• Online Access: https://dx.doi.org/10.5281/zenodo.5541656; https://zenodo.org/record/5541656

Diplura macrura (C.L. Koch, 1841) Figs 1–13, 26 Mygale macrura C.L. Koch, 1841: 38, fig. 715 (♂). Diplura bicolor Simon, 1889: 215 (♀) syn. nov. Thalerothele uniformis Mello-Leitão, 1923: 105, fig. 4 (♂) syn. nov. Thalerothele minensis Mello-Leitão, 1926: 314, figs 1-3 (♂) syn. nov. Thalerothele aurantiaca Mello-Leitão, 1943: 255 (♀) syn. nov. Diplura bicolor – Mello-Leitão 1937: 3, fig. 3 (♂). — Bücherl 1957: 385, figs 12, 12a. Diplura macrura – C.L. Koch 1850: 75. — Raven 1985: 74, figs 18–23. Thalerothele uniformis – Bücherl, Timotheo & Lucas 1971: 128, figs 12–14 (syn. T. minensis, T. aurantiaca ). Diplura uniformis – Platnick 1993: 89 (transfer). — Silva-Moreira et al . 2010: 32. Linothele bicolor – Platnick 1998: 120 (transfer). Diagnosis Both sexes of this species have a strongly contrasting color pattern, with a reddish brown carapace and a dark brown abdomen, shared only with Diplura paraguayensis (Gerschman & Schiapelli, 1942). In mature D. macrura , the abdomen is uniformly dark brown, without spots or any visible markings. However, according to its original description (Gerschman & Schiapelli 1942), D. paraguayensis has a blackish brown reticulate on abdomen dorsum. Moreover, the lyra of D. paraguayensis has 13 setae (Gerschman & Schiapelli 1942: pl. ix; Schiapelli & Gerschman 1968, fig. 7), in contrast with the 7–9 setae in D. macrura . The bulb of D. macrura (Figs 6–7) has an embolic base much larger than in D. paraguayensis (Schiapelli & Gerschman 1968: figs 14–15). Also, the embolus is about 2× longer than the bulb in D. macrura and 3× longer in D. paraguayensis . The spermathecae of D . macrura has a longer and thinner stalk (Fig. 13) than the females of D . paraguayensis from Argentina (Goloboff 1982: 1). Also, the distal lobes are larger and spread over the distal third of the stalk in D . macrura , while they are smaller and concentrated in the apex in D . paraguayensis . Type material Mygale macrura : BRAZIL: ♂, holotype, Minas Gerais, São João del Rei, [1830], Sellow, Friedrich W. (MfN 2083), examined. Diplura bicolor : BRAZIL: 2 ♀♀, syntypes, Minas Gerais, Catas Altas, Caraça (coll. E. Simon, MNHN AR 4932 B337), examined (photos). Thalerothele uniformis : BRAZIL: ♂, holotype, São Paulo, E. of Garbe (MZSP?, MNRJ?), not located. Thalerothele minensis : BRAZIL: ♂, holotype, Minas Gerais, Ouro Preto, Magalhães Gomes (Mello- Leitão collection 880, MNRJ 1360), not located. Thalerothele aurantiaca : BRAZIL: ♀, holotype, Minas Gerais, Ouro Preto/Mariana: Itacolomi, O. Leonardos (MNRJ 53945), not located. Material examined BRAZIL: 1 ♂, Minas Gerais, Parque Nacional Serra do Cipó, R. Bertani, R. Martins, C.S. Fukushima & M.P. Pavani leg. (MNRJ); 1 ♀, 2 juvs, Caeté, 11–19 Feb. 1961, P. Vanzolini, H. Britski & N. Menezes leg. (MZSP 6576); 1 ♂, Catas Altas, Reserva Particular de Patrimônio Natural Serra do Caraça, 24 Apr.– 1 May 2002, Equipe Biota (IBSP 13148); 2 ♂♂, 1 juv., Lima Duarte, Parque Estadual de Ibitipoca, 21 Mar. 1997, A. Oliveira & B. Souza leg. (IBSP 8390); 1 ♂, Parque Estadual de Ibitipoca, Apr. 1997, A. Oliveira & B. Souza leg. (IBSP 8432); 1 ♀, juv., Ouro Preto, Estação Ecológica de Tripuí, R. Bertani, R. Martins, C.S. Fukushima & M.P. Pavani leg. (MNRJ 4467 and MNRJ 4490, respectively); 1 ♀, Estação Ecológica de Tripuí, 1220 m, 20°22'92.7" S, 43°33'29.3" W, R. Bertani, R. Martins, C.S. Fukushima & M.P. Pavani leg. (MNRJ 4502); 1 ♂, Estação Ecológica de Tripuí, 1236 m, 20°22'93.2" S, 43°33'28.7" W, R. Bertani, R. Martins, C.S. Fukushima & M.P. Pavani leg. (MNRJ 4496); 1 ♀ (MNRJ 4487), 1 ♂ (MNRJ 4497), 1 ♀ (MNRJ), Estação Ecológica de Tripuí, 1246 m, 20°22'93.2" S, 43°33'28.7" W, R. Bertani, R. Martins, C.S. Fukushima & M.P. Pavani leg.; 1 ♀, Estação Ecológica de Tripuí, 20°22'6.62" S, 43°32'5.74" W, R. Bertani, R. Martins, C.S. Fukushima & M.P. Pavani leg. (MNRJ 4509); 1 juv., Estação Ecológica de Tripuí, 20°22'9.70" S, 43°33'1.04" W, R. Bertani, R. Martins, C.S. Fukushima & M.P. Pavani leg. (MNRJ 4498); 1 ♀, Estação Ecológica de Tripuí, 20°22'915" S, 43°33'227" W, R. Bertani, R. Martins, C.S. Fukushima & M.P. Pavani leg. (MNRJ 4489); 1 ♂, Estação Ecológica de Tripuí, 20°22'93.2" S, 43°33'26.7" W, R. Bertani, R. Martins, C.S. Fukushima & M.P. Pavani leg. (MNRJ 4505); 1 ♂, Estação Ecológica de Tripuí, 20°22'93.2" S, 43°33'28.7" W, R. Bertani, R. Martins, C.S. Fukushima & M.P. Pavani leg. (MNRJ 4479); 1 juv., Estação Ecológica de Tripuí, 20°22'93.2" S, 43°33'28.7" W, R. Bertani, R. Martins, C.S. Fukushima & M.P. Pavani leg. (MNRJ); 1 juv., Estação Ecológica de Tripuí, 20°22'970" S, 43°33'104" W, R. Bertani, R. Martins, C.S. Fukushima & M.P. Pavani leg. (MNRJ 4492); 1 ♂, Estação Ecológica de Tripuí, 20°22'92,2" S, 43°33'13" W, 8 Jan. 2006, R. Bertani, R. Martins, C.S. Fukushima & M.P. Pavani leg. (MNRJ); 1 ♀, Estação Ecológica de Tripuí, 20°22'93.2" S, 43°33'287" W, 7 Jan. 2005, R. Bertani, R. Martins, C.S. Fukushima & M.P. Pavani M. leg. (MNRJ 4333); Ouro Preto: 1 juv., Parque Estadual do Itacolomi, R. Bertani, R. Martins, C.S. Fukushima & M.P. Pavani leg. (MNRJ); 1 juv., Parque Estadual do Itacolomi, R. Bertani, R. Martins, C.S. Fukushima & M.P. Pavani leg. (MNRJ 4515); 1 ♂, Parque Estadual do Itacolomi, 1,306 m, 20°22'662" S, 43°32'574" W, R. Bertani, R. Martins, C.S. Fukushima & M.P. Pavani leg. (MNRJ 4495); 1 ♀, Parque Estadual do Itacolomi, 20°25'638" S, 43°30'399" W, R. Bertani, R. Martins, C.S. Fukushima & M.P. Pavani leg. (MNRJ 4507). Description Male (MNRJ 4496) (Figs 3–9) Carapace: 6.4 long, 5.0 wide. Abdomen: 6.6 long. Spinnerets: PMS 1.3 long; PLS, total length 6.8, basal article 2.0, middle 2.2, distal 2.6, respectively. Legs: see Table 1. Carapace: length/width 1.3; flat, cephalic area slightly raised, thoracic furrows shallow and wide. Fovea: short, deep, recurved. Carapace covered with short, thin setae, interspersed with some longer and thicker setae; border with abundant long and thick setae pointing outwards, increasing in number towards posterior angles. Clypeus almost totally hidden by the bulging eye tubercle, frontal margin bearing 5 thick, long, erect setae. Eye tubercle: 0.6 long, 1.0 wide, area between posterior eyes covered with thin setae and bearing 4 thicker, longer setae. AME 0.3, almost spherical, but a bit longer than wide, set apart by 0.7× their diameter. ALE elliptical, much longer than wide, its length about 0.6× the AME diameter. PME small, with flattened lens, longer than wide, its length about 0.5× AME diameter. PLE elliptical, much longer than wide, its length a bit less than 0.7× AME diameter. PME and PLE clearly set apart by around 0.4× the PME length. Anterior eye row slightly recurved, posterior eye row recurved. Eye rows with similar width. Chelicerae: promargin with 11 teeth on left and 9 on right chelicera. Plectrum with 5 thick, long setae. Labium: length/width 0.8, no cuspules. Labio-sternal groove deep with elongated sigilla. Sternum: about 45% longer than wide, very similar to the sternum of D. lineata . Posterior angle in a blunt point, not separating coxae IV. Sigilla: three pairs, spherical, with a subtle increase in size from anterior to posterior, all near margin. Palp (Fig. 5): relatively short, without retrolateral spines, femur: d1–2–0, pl0–0–1, tibia pl 0–1–0, v1–2 –0. Tibia: length 2.3, width 0.9, short, incrassated, thinner at the basis and apex. Maxillae: length/width 1.5. Cuspules: 13 spread over ventral inner heel. Lyra at the ventral side of the maxilla, formed by 8 modified thick, long setae, increasing in size from basis to apex of the lyra, weakly curved at apical portion. Legs: Leg formula 4123. Legs covered with short, thin, horizontal black setae and with some longer, thicker, erect black setae. All tarsi with thin scopula, throughout the length of the article, divided by two series of thicker setae at the middle line of the ventral face. Metatarsus I with undivided thin scopula, covering the distal half of the retrolateral side. Metatarsus II similar to I, but with scopula on both sides. Metatarsi III–IV without scopula. All tarsi provided with numerous small cracks covering almost all the ventral and lateral faces, except by the basis and tip of the article. Leg I (Figs 3–4): tibia I with a relatively short distal retrolateral spur, curved and blunt, placed at its ventral corner. Megaspine pointed, slightly sinuous, especially on the apex, almost 2x longer than the spur. Metatarsus I with a distinct retrolateral tubercle placed ventrally at the beginning of the median third, conical, pointed and facing towards the tip of article. Ventrally, there is one spine near the apex and another one on the median third, placed much ahead of the tubercle. Fringe formed by many thick and long spiniform setae and spines (clasper) covering almost all prolateral side of the metatarsus (Fig. 4). Spines: leg I: femur d1–2–0, pld0–0–1, rld0–0–1, patella p0–0–1 left, p0 right, r0 left, 0–1–0 right, tibia p0–2–0, v0–1–1ap (apophysis), metatarsus p0–2–0, v0–1–1ap; leg II: femur d1–2–0, pld1–0–1 left, pld0–0–1 right, rld0–0–1 left, rld0 right, patella p0–1–1 left, p0–0–1 right, tibia p0–2–0, v1–1 –2ap left, v1 –0–2ap right, metatarsus p1–1–1 left, p1–1–0 right, v1–2 –2ap left, v1–2 –1ap; leg III: femur d1–2–0, pld0 left, pld0–1–1 right, rld0–1–1 left, rld0–2–1 right, patella p0–1–1 left, p0–1–1 right, r0–1–0 left, r1–1–0 right, tibia p0–2–0 left, p0–2–1 right, r1–1–1, v1–1 –2ap; metatarsus p2–2–1 left, p0–4–1 right, r1–1–1 left, r1–2–1 right, v0–4–3ap left, v0–3–3ap right; leg IV: femur d2–1–0 left, d1–2–0 right, rld0– 0–1 left, rld0–0–2 right, patella r0–1–0, tibia p0–1–0 left, p0–2–0 right, r1–2–1 left, r1–1–1 right, v2– 1 –2ap left, v1 –0–2ap right, metatarsus d1–0–0 left, d1–1–0 right, p1–2–1, r1–2–1, v1–5 –3ap. Claws: ITC without teeth. Teeth at STC: all claws with a small spur at the basis in both sides leg I: inner 6–7 and outer rows 5–6; leg II: inner row 9, outer row 8–9; leg III: inner row 6–7, outer row 6; leg IV: inner and outer rows 6–7. Bulb (Figs 6–9): piriform and conical in a retrolateral view, with length about ¾ of its width. Embolus about 2× longer than the bulb itself, with a wide base in relation to the globose part of the bulb and tapering towards the apex. Embolus clearly curved since its base and tapering from the basal third to the apex, clearly seen both on retrolateral and prolateral views. However, in dorsal view, the embolus is just slanted, almost straight. On ventral view, spermatic duct large at the base, tapering regularly towards the tip, but much thinner from the distal third on. Female (MNRJ 4487) (Figs 2, 10–13) Total length 15.3. Carapace: 7.3 long, 5.8 wide. Abdomen: 8.0 long, 5.0 wide. Spinnerets: PMS 1.4 long; PLS, total length 5.9, basal article 2.3, middle 1.6, distal 2.0, respectively. Legs: see Table 2. Females are very similar to males except by its bigger size and the following characteristics: carapace length/ width 1.3. Clypeus very narrow, with the frontal margin carrying five thick, long setae, facing forward. Eye tubercle with a thick seta on its anterior margin. Area between eyes with two longer and thicker setae. AME separated by around their diameter. PME and PLE less separated. Chelicera with 10–11 promarginal teeth, on the right and left chelicera, respectively. Plectrum with 6 thick, long setae. Maxilla with 16 (left) or 19 (right) cuspules. Lyra (Figs 10–11) as in male, formed by 7 modified setae. Tarsi I–II with scopula similar to male, but tarsus III with a very thin scopula and tarsus IV without scopula. Metatarsi I–II with undivided thin scopula, covering both sides and the ventral face. All tarsi provided with few small cracks covering only the median area of the ventral and lateral faces. Spines: leg I: femur d1–2–0, pld0–0–1; patella 0; tibia p0–0–1, v0–0–2ap (apophysis); metatarsus v0–3–2ap; leg II: femur d1–1–0, pld0–0–1; patella p0–0–1; tibia p0–0–1, v0–0–2ap; metatarsus v0–3–2ap; leg III: femur d1–0–0, rld0–0–2 left, rld0–0–1 right; patella p0–1–1 left, r0–1–0; tibia p1–1–0 left, p0–2–0 right, r0–2–0, v0–0–1ap left, v0–0–2ap right; metatarsus p1–2–1 left, p1–3–1 right, r1–1–1, v0–4–3ap; leg IV: femur d1–1–0 left, d1–2–0 right, rld0–0–1; patella r0–1–0; tibia p1–1–0, r2–1–0, v0–0–1ap left, v–0– 0–2ap right; metatarsus p0–2–1 left, p1–2–1 right, r2–1–2 left, r1–1–2 right, v1–3 –2ap. Spermathecae (Fig. 13): separated by about 70% of its length, with a thick stem, keeping a similar width up to the distal lobules. Stem curved forward from the median part on, bearing three large apical lobules of variable sizes. Color pattern Both sexes with carapace reddish brown, covered by abundant setae of a weakly coppery hue. Thoracic furrows and cephalic area darker. Eye area black. Legs, palps, chelicera and spinnerets dark brown, with a clear contrast to the color of carapace. Labium, sternum and leg coxae orange, with darker sigilla. Dorsum of the abdomen uniformly dark brown without markings or stripes, venter usually lighter colored than dorsum, but with a dark hue in some males. Variation Chelicera with 8–11 promarginal teeth. Lyra at the ventral side of the maxilla, formed by 7 to 9 modified setae (Figs 10–12). The maxillary cuspules may vary from 13 to 17 in males and from 16 to 19 in females. Synonymy and notes When describing Mygale macrura , C.L. Koch (1841: 39) indicated that the type-locality of the species was “WestIndien, St. Juan”. Nine years later, he transferred the species to his new genus Diplura , without mentioning the type-locality (C.L. Koch 1850: 75). The type-locality indicated by Koch 1841 was cited differently by subsequent authors, for example, “Westindien” (Ausserer 1871: 178), “St. Jean, Antilles” (Simon 1903: 963), “West Indies” (Petrunkevitch 1911: 60), or “Saint John, U.S. Virgin Islands” (Muchmore 1993: 32). All those subsequent authors did not mention any additional specimen of D. macrura and were certainly restating the information given by C.L. Koch (1841). Banks (1909: 155) was the first arachnologist to record D. macrura in Cuba, from Pinar Del Rio, as “ Ischnothele macrura Koch ”, but he made no comments on the implicit genus transfer, nor did he include any description of the specimens he examined. Lutz (1915: 77) and Petrunkevitch (1926: 27) also pointed out that D. macrura had previously been recorded from Cuba. While Lutz expressly cited Banks (1909), Petrunkevitch did not mention its source. However, he was probably also citing Banks (1909), as he stated that “ Diplura macrura C. Koch, has been also reported from Cuba”. In his paper on endemic spiders from Cuba, Alayón (2000: 38) also included D. macrura without any further comment. Currently, the only record for D. macrura included in the World Spider Catalog (2015) is Cuba, without any mention of Koch’s type-locality. In her paper on Cuban spiders though, Bryant (1940: 260) considered Banks’ record a misidentification of Ischnothele longicauda Franganillo, 1936. She based her opinion on the relative abundance of the latter species in Cuba. The family Dipluridae is scarcely represented in Cuba, with only I. longicauda and two species of Masteria L. Koch, 1873 recorded for the country (World Spider Catalog 2015), not considering the erroneous D. macrura citation by Banks. Therefore, the only specimen of D. macrura undoubtedly cited for Cuba, West Indies or any other Caribbean locality is the holotype. Surprisingly, an analysis of the holotype of D. macrura indicates that the type-locality was misinterpreted by Koch. The holotype label clearly indicates “Koch *. typ. S. João d. Rey. Sello” (Fig. 1, right). On the other hand, there is an additional label (Fig. 1, left) inside the vial that contains the type indicating “ Macrura N. Koch. * fg. 715. Cametá Sieber”, referring to Cametá, a locality in the state of Pará, northern Brazil. This additional label was probably wrongly added later in the vial, as there are no records for D. macrura from Pará. Up to now, all the specimens of D. macrura we examined came only from the state of Minas Gerais. Therefore, it is clear that the type-locality is really São João del Rei, central Minas Gerais state, Brazil. Friedrich Sellow (or Sello, 1789–1831), a famous Prussian naturalist, collected a large number of zoological specimens in Minas Gerais state, such as insects (Papavero 1973) and birds (Rego et al. 2013). Furthermore, most of Sellow´s specimens are also deposited at MfN. Diplura bicolor was described from Caraça, Minas Gerais State, Brazil, based on two female syntypes (Simon 1889: 215). It is, currently, placed in the genus Linothele , although it was considered a Diplura by former authors (Mello-Leitão : Published as part of Denis Rafael Pedroso, Pedro de Souza Castanheira & Renner Luiz Cerqueira Baptista, 2016, Redescription and synonymies of Diplura macrura (C. L. Koch, 1841) and D. lineata (Lucas, 1857), with notes on the genus (Araneae, Dipluridae), pp. 1-21 in European Journal of Taxonomy 210 on pages 3-11, DOI: 10.5852/ejt.2016.210, http://zenodo.org/record/269325 : {"references": ["Koch C. L. 1841. Die Arachniden. Vol. 9. C. H. Zeh'schen Buchhandlung, Nurnberg", "Simon E. 1889. Arachnides. In: Voyage de M. E. Simon au Venezuela (Decembre 1887 - Avril 1888). 4 e Memoire. Annales de la Societe Entomologique de France (6) 9: 169 - 220. 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