Chaoborus albipes Salmela & Härmä & Taylor 2021, stat. rev.
Chaoborus albipes (Johannsen, 1903), stat. rev. Corethra albipes Johannsen 1903: 398 (original description of female holotype), wing plate 39 fig. 11. Richardson 1912: 202 (redescription of larva, especially in comparison to C. plumicornis var. americana (= Chaoborus americanus ), description of mal...
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2021
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| Online Access: | https://dx.doi.org/10.5281/zenodo.4564950 https://zenodo.org/record/4564950 |
| Summary: | Chaoborus albipes (Johannsen, 1903), stat. rev. Corethra albipes Johannsen 1903: 398 (original description of female holotype), wing plate 39 fig. 11. Richardson 1912: 202 (redescription of larva, especially in comparison to C. plumicornis var. americana (= Chaoborus americanus ), description of male, short comments on pupa and adult female, biology), larval antennae fig. 1, male wing fig. 2.1, female wing fig. 2.2 Sayomyia albipes : Felt 1904: 363 (in part, description of female copied from Johannsen 1903, description of pupa and male). Felt 1905: 497 (verbal description of male hypopygium). Dyar 1905: 16 (key to larvae, confused with C. punctipennis ). Chaoborus (Chaoborus) albipes : Dyar & Shannon 1924: 211 (comparison to Chaoborus crystallina and C. flavicans , distribution in USA). Hennig 1968: 74 (list of Nearctic species, literature of larvae and pupae). Chaoborus albipes : Johannsen 1934: 44 (identification key, diagnoses of larva and pupa, var.a. consists of larvae with 25 or more anal fan setae and somewhat broader labral blades), larva habitus plate 43 fig. 166, pupa habitus fig. 167, labral blades fig. 171. Dickinson 1944: 357 (verbal description of adult and larva, perhaps based on earlier references, occurrence in Wisconsin), labral blade fig. 229. Belkin et al. 1966: 22 (location of holotype, type locality, bionomics, as a syn. of C. flavicans ). Sayomyia rotundifolia Felt 1904: 366 syn. nov. (description of female, male, larva and pupa), female wing plate 13 fig. 2, male wing plate 13 fig. 3, male hypopygium plate 40 fig. 2. Felt 1905: 497 (verbal description of male hypopygium). Dyar 1905: 16 (key to larvae). Chaoborus rotundifolia : Johannsen 1934: 43 (identification key, following the concept by Felt). Chaoborus (Chaoborus) rotundifolia : Hennig 1968: 74 (list of Nearctic species, literature of larvae and pupae). Chaoborus crystallina : Matheson 1925: 159 ( Sayomyia rotundifolia as a new synonym of C. crystallina ). Chaoborus crystallinus : Yamada 1932: 230 (redescription of female), habitus photo of adult female on page 230. Matheson 1944: 95 (identification key), male hypopygium plate 10 fig. 4. Chaoborus (Chaoborus) flavicans : Cook 1956: 23, in part. (redescription of C. (C.) flavicans includes both C. flavicans and C. albipes ), parameres (fig. 16H). Chaoborus flavicans : Strickman 1980: (in vitro copulation and oviposition), egg mass. fig. 1. Likely the species involved here is C. albipes , because material studied was from a “shallow, woodland pond”. Ogawa 2007: (phylogeny), mandible fig. 2.15b, parameres fig. 2.51b. Luoto & Nevalainen 2009: in part. (paleolimnology), mandibles in fig. 2a represent C. albipes . Chaoborus cf. flavicans : Dupuis et al. 2008: (molecular analysis, phylogeny, distribution, ecology, morphology), mandible fig. 2A. An et al. 2010: (molecular phylogeny, partly based on same material as in Dupuis et a. 2008). Taylor et al. 2015: (distribution, ecology). Ballinger et al. 2014: (RNA virus infection, distribution). Ballinger et al. 2017: (RNA virus infection, distribution). Material examined. Type material. Holotype. [“O.A.J. det 1759” “ Corethra albipes Jo. Ithaca N.Y. 1901 HOLOTYPE no 2968 (partly hand written)” “Cornell University Dept. of Entomology”] white label on slide. [“HOLOTYPE Cornell U. No. 2968”] red label on slide (CUI). An additional white label is glued on left upper corner, [Chaoborus flavicans (Meig.) det. E.F. Cook 1951]. The holotype of C. albipes consists of a slide mounted wing only, the rest of the specimen is lost (J.J. Dombroskie, pers.comm.). The holotype was studied from photos. Lectotype of Sayomyia rotundifolia , by present designation (J. Salmela, Fig. 2a,b,d). [“Entomologic division (printed)” “ Sayomyia rotundifolia ³” “Karner N.Y.” “Aug. 5 ‘04” “Genitalia in Bal. (hand written)” “N.Y. state museum (printed)”]; lectotype of S. rotundifolia consists of a slide mounted male abdominal terminalia. Paralectotypes, by present designation. 1 male, [“Entomologic division (printed)” “ Sayomyia rotundifolia ” “Karner N.Y.” “1 July 04” “Wing in Bal. ³ s.no. 14 (hand written)” “N.Y. state museum (printed)”]; a wing on a slide (Fig. 2e). 1 larva, [“Entomologic division (printed)” “ Sayomyia rotundifolia ” “Karner N.Y.” “18 July ‘04” “Larva in Bal. (hand written)” “N.Y. state museum (printed)” (Fig. 2c,f); one slide mounted larva, intact, i.e. mouthparts etc. not detached; most likely belong to C. flavicans , not C. albipes . 1 female, [“Entomologic division (printed)” “ Sayomyia rotundifolia ” “Karner N.Y.” “Aug. 2 04” “Wing in Bal. ♀ (hand written)” “S.no.14 (handwritten)” “N.Y. state museum (printed)”]; one slide-mounted female wing (Fig. 2g). According to the original description Felt studied only one male specimen, but different dates on the labels indicate that lectotype male and paralectotype male are different specimens. Other material. Finland. Ab: Salo, Härjänsilmä, Salmela J. leg. 2. V.2020, 6 larvae, NVO.LMM-el-20-5; 1 male e.l., NVO.LMM-el-20-15; 1 male e.l., NVO.LMM-el-20-16; 1 male e.l., NVO.LMM-el-20-17; 1 male e.l., NVO.LMM-el-20-18; 1 male e.l., NVO.LMM-el-20-23; 1 female e.l., NVO.LMM-el-20-27; 1 male e.l., NVO. LMM-el-20-28; 1 male e.l., NVO.LMM-el-20-29; 1 male pupa, e.l., NVO.LMM-el-20-34; 4 males e.p., NVO. LMM-el-20-49; 2 males e.p., NVO.LMM-el-20-55 (LMM); 1 male e.p. (ABC). N: Sipoo, Jöusjärv, Salmela J. leg. 2. V.2020, 1 pupa e.l., NVO.LMM-el-20-2; 1 male e.l., NVO.LMM-el-20-19; 1 male e.l., NVO.LMM-el-20-20; 1 male e.l., NVO.LMM-el-20-21; 1 male e.p., NVO.LMM-el-20-31; 1 male e.p., NVO.LMM-el-20-47; 1 female e.p., NVO.LMM-el-20-54 (LMM). Sipoo, Fallträsk, Salmela J. leg. 2. V.2020, 1 larval and pupal exuviae on slide, adult escaped (LMM). Ta: Tammela, Kärjensuo 1, Härmä O. leg. 9.IV.2020, 1 male e.l., LG.6092; 1 male e.l., LG.6092; 1 male e.l., LG.6093; 1 male e.l., LG.6094; 1 male e.l., LG.6109; 1 pupa e.l., LG.6110; 1 female e.l., LG.6111; 1 male e.l., LG.6128; 1 male e.l., LG.6129; 1 male e.l., LG.6130; 1 male e.l., LG.6132; 1 male e.l., LG.6164; 1 female e.l., LG.6165; 6 larva, LG.6165 (FLHM). Tammela, Kärjensuo 2, Härmä O. leg. 9.IV.2020, 1 larva (FLHM). Tammela, Ammeenpohja, Härmä O. leg. 16.IV.2020, 2 larvae, LG.6113; 1 male e.l., LG.6227; 1 male e.l., LG.6228; 1 female e.l., LG.6229; 1 pupa e.l., LG.6230; 1 male e.l., LG.6231; 1 female e.l., LG.6232; 1 pupa, LG.6234; 1 male e.l., LG.6246 (FLHM). Tammela, Lehmälammi, Härmä O. leg. 21.IV.2020, 9 larvae, LG.6193; 1 male e.l., LG.6236; 1 male e.l., LG.6237; 1 female e.l., LG.6238; 1 female e.l., LG.6239; 1 male e.l., LG.6240; 1 female e.l., LG.6241; 1 male e.l., LG.6242; 1 female e.l., LG.6252; 1 pupa e.l., LG.6259; 1 pupa e.l., LG.6260 (FLHM). Tammela, Paskolammi, Härmä O. leg. 30.IV.2020, 5 larvae, LG.6262 (FLHM). Tammela, Vehkala, Härmä O. leg. 21.IV.2020, 10 larvae, LG.6198; 1 female e.l., LG.6243; 1 female e.l., LG.6244; 1 female e.l., LG.6245; 1 female e.l., LG.6253; 1 male e.l., LG.6255; 1 female e.l., LG.6256; 1 female e.l., LG.6257; 1 pupa e.l., LG.6258 (FLHM). Jokioinen, Kaitalammi, Härmä O. leg. 13.IV.2020, 10 larva, LG.6126; 5 larvae, LG.6147; 1 male e.l., LG.6169; 1 male e.l., LG.6170; 1 male e.l., LG.6172; 1 male e.l., LG.6173; 1 female e.l., LG.6173; 1 female e.l., LG.6175; 1 male e.l., LG.6176; 1 male e.l., LG.6177; 1 male e.l., LG.6178; 1 male e.l., LG.6179; 1 male e.l., LG.6180; 1 female e.l., LG.6181; 1 pupa e.l., LG.6182; 9 males e.l., LG.6183; 8 females e.l., LG.6184; 1 male e.l., LG.6202; 1 female e.l., LG.6203; 1 female e.l., LG.6204; 1 male e.l., LG.6205; 1 pupa e.l., LG.6226; 1 pupa e.l., LG.6248; 1 pupa e.l., LG.6249; 1 pupa e.l., LG.6250 (FLHM). Somero, Hossinoja, Härmä O. leg. 10.IV.2020, 1 male e.l., LG.6102; 1 male e.l., LG.6103; 1 male e.l., LG.6104; 1 male e.l., LG.6105; 1 female e.l., LG.6106; 1 female e.l., LG.6107; 1 male e.l., LG.6108; 1 male e.l., LG.6122; 1 male e.l., LG.6124; 4 larva, LG.6125; 1 male e.l., LG.6131; 1 female e.l., LG.6133; 1 pupa e.l., LG.6134; 1 pupa e.l., LG.6166 (FLHM). Somero, Jorri, Härmä O. leg. 18.IV.2020, 2 larvae, LG.6163 (FLHM). Somero, Salakkajärvi W, Salmela J. leg. 2. V.2020, 4 larvae, NVO.LMM-el-20-9; 1 male e.l., NVO.LMM-el-20- 12; 1 male e.l., NVO.LMM-el-20-13; 1 male e.l., NVO.LMM-el-20-14; 1 pupa e.l., NVO.LMM-el-20-25; 1 male e.l., NVO.LMM-el-20-26; 5 male e.p., NVO.LMM-el-20-32; 2 male e.p., NVO.LMM-el-20-50; 1 male e.p., NVO. LMM-el-20-56 (LMM). Somero, Salakkajärvi W, Härmä O. leg. 10.IV.2020, 1 male e.l., LG.6095; 1 larva, LG.6114 (FLHM). Somero, Äijämö, Härmä O. leg. 18.IV.2020, 1 larva, LG.6156; 1 larva, LG.6158; 1 male e.l., LG.6212; 1 female e.l., LG.6213; 1 male e.l., LG.6214; 1 male e.l., LG.6215; 1 male e.l., LG.6217; 1 pupa e.l., LG.6218; 1 pupa e.l., LG.6219; 1 pupa e.l., LG.6220; 1 pupa e.l., LG.6221; 1 pupa e.l., LG.6222; 1 pupa e.l., LG.6223 (FLHM). Kouvola, Konttisuo, Salmela J. leg. 6. V.2019, 1 larva, head, tail, respiratory organs of prepupa (taken beneath larval skin) on slide, BOLD, NVO.CUL-2019-26 (LMM); Kymijoen vesi ja ympäristö leg. 18.XI.2015, 1 larva, head & tail on slide, BOLD, NVO.ins2018-783 (LMM); 1 larva, head + tail on slide, NVO.CUL-2019-84 (LMM). Sa: Savonlinna, Ukonlampi, Salmela J. leg. 07. V.2019, 1 male e.l., pinned, larval and pupal exuviae on slide, hypopygium on slide, BOLD, NVO.LMM-el-15; 1 male e.l., pinned, NVO.LMM-el-17; 1 female e.l., larval and pupal exuviae on slide, NVO.LMM-el-42; 1 male e.l., pinned, NVO.LMM-el-57; 1 male e.l., pinned, pupal exuviae on slide, BOLD, NVO.LMM-el-61; 3 males e.p., NVO.LMM-el-64; 1 male e.l., pinned, pupal exuviae and hypopygium on slide, BOLD, NVO.LMM-el-76; 1 male e.l., pinned, NVO.LMM-el-99 (LMM). Sb: Joroinen, Ulminmäki, Salmela J. leg. 07. V.2019, 1 male e.l., pinned, larval and pupal exuviae on slide, hypopygium on slide, BOLD, NVO.LMMel-10; 1 male e.l., pinned, larval and pupal exuviae on slide, hypopygium on slide, BOLD, NVO.LMM-el-11; 1 male e.l., pinned, larval and pupal exuviae on slide, hypopygium on slide, NVO.LMM-el-13; 1 male e.l., larval and pupal exuviae on slide, hypopygium on slide, NVO.LMM-el-29; 1 female e.p., pinned, pupal exuviae on slide, NVO. LMM-el-39; 1 male e.p., NVO. LMM-el-40; 1 male e.l., pinned, pupal exuviae on slide, NVO.LMM-el-41; 3 males e.p., parameres of two specimens glued on slide, NVO.LMM-el-49; 1 male e.p., pupal exuviae, wings, legs, head, hypopygium on slide, NVO.LMM-el-50 (LMM). Tb: Toivakka, Kataislammit, Salmela J. leg. 1. V.2020, 1 larva, NVO.LMM-el-20-1; 1 male e.l., NVO.LMM-el-20-42; 1 male e.l., NVO.LMM-el-20-43; 1 pupa e.l., NVO.LMMel-20-44; 1 male e.l., NVO.LMM-el-20-45 (LMM). Obb: Rovaniemi, Pietarinlampi, Salmela J. leg. 5. VI.2020, 1 male e.p., pupal exuviae and hypopygium on slide, NVO.LMM-el-20-105; 1 male e.p., adult pinned, pupa exuviae on slide, NVO.LMM-el-20-107; 1 male e.p., NVO.LMM-el-20-108 (LMM). Lkoc: Kittilä, Muotrikkilehto, Salmela J. leg. 27.VII.2019, 1 larva, NVO.LMM-el-608 (LMM); 1 larva, NVO.LMM-el-610 (LMM); 1 larva, head & tail on slide, BOLD, NVO.CUL-2019-78 (LMM); 1 larva, head & tail on slide, BOLD, NVO.CUL-2019-79 (LMM). Kittilä, Papinmutka, Salmela J. leg. 28.VII.2019, 1 female, pupal exuviae and female in EtOH, BOLD, NVO.LMMel-526 (LMM). Lkor: Savukoski, Rouvoivanselkä, Salmela J. leg 05. VI.2019, 1 male e.l., larval and pupal exuviae, hypopygium on slide, NVO.LMM-el-206; 1 male e.l., NVO.LMM-el-211; 1 male e.l., larval and pupal exuviae on slide, legs, wings, head on slide, torso EtOH, NVO.LMM-el-212; 1 male e.l., larval and pupal exuviae on slide, legs, wings, head on slide, torso EtOH, NVO.LMM-el-213; 1 male e.l., larval and pupal exuviae on slide, legs, wings, head, hypoopgium on slide, torso EtOH, NVO.LMM-el-214; 1 male e.l., NVO.LMM-el-215; 1 female e.l. pinned, larval & pupal exuviae on slide, NVO.LMM-el-216; 1 male e.l., NVO.LMM-el-217; 1 male e.l., NVO.LMM-el- 218; 1 male e.l., NVO.LMM-el-219; 1 male e.l., NVO.LMM-el-220; 1 female e.l., NVO.LMM-el-221; 1 male e.l., NVO.LMM-el-222; 1 male e.l., LMM-el-223; 1 male e.l., NVO.LMM-el-224; 1 male e.l., NVO.LMM-el-225; 1 male e.l., NVO.LMM-el-226; 1 male e.l., NVO.LMM-el-227; 1 pupa e.l., NVO.LMM-el-228; 1 pupa e.l., NVO. LMM-el-229; 1 male e.l., NVO.LMM-el-230; 1 male e.l., NVO.LMM-el-231; 1 female e.l., NVO.LMM-el-232; 1 female e.l., larval and pupal exuviae on slide, NVO.LMM-el-233; 1 pupa e.l., NVO.LMM-el-234; 1 male e.l., NVO. LMM-el-290; Laine E. & Salmela J. leg. 13. VI.2018, 5 larvae 5 pupae, one teneral hypopogium on slide, NVO. ins2018-420; 10 larvae 3 pupae, one teneral hypopogium on slide NVO.ins2018-421; 1 pupa teneral hypopogium on slide, NVO.ins2018-427; 1 pupa, BOLD, NVO.ins2018-624; 1 pupa, BOLD, NVO.ins2018-625; 1 larva, BOLD, NVO.ins2018-626; 1 larva, BOLD; NVO.ins2018-627; 1 larva, BOLD, NVO.ins2018-628; 1 larva, NVO.ins2018- 681; Laine E. leg. 12.VII.2018, 7 larvae I and II instar, NVO.ins2019-195 (LMM). Norway. Buskerud, Kongsberg, Karlshaug - N Svensketjern, 59.5302 9.5642, 495 masl, K.M. Olsen leg. 31. V.- 26. VI.2013, 20 males, 1 on slide, J.nr. BAB 470603 (KMO). Russia. South Urals, Osipovka, Salokannel, J. leg. 05.VIII.2018, 1 female, BOLD, NVO.ins2018-457 (LMM). Primorskiy Kray, Kiparisovo (Tazyozhnyi), 43.4858 131.975, E. Zakharov leg. 7.VII.2014, 1 female, BOLD, BIOUG 15361 - C01 (CBG) Japan. Tokyo, Yotsuya, 35.68 139.72, leg? 1948, 3 males 1 female in EtOH (NMNS). Sado Island, 38.03 138.33, I. Tosayabashi leg. 16. VI.1917, 1 male, pinned hypopygium in microvial (NMNS). Shiba, Tokio, 35.65 139.74, S. Yamada leg. 20.VIII.1919, 1 male, pinned (NMNS). Hokkaido, K. Tanaka leg., no date, 3 males, pinned (two hypopygia in microvial), A-0213, (NMNS). Matsuyama-shi, Higashino, 33.8411 132.8111, J. Oku leg. 19. V.2016, 1 male, pinned, hypopygium in microvial (ELEU). Honsu, Niigata, Matsunoyama-Kannonji, Kato D. leg. 13.VIII.2020, 1 male, NVO. JAP-06; 1 male, NVO. JAP-07; 1 male, NVO. JAP-09; 1 male, NVO.JAP-11 (LMM). Canada. Ontario, Roseneath L., 44.19 -78.05, H.G. James leg. 1963, 1 male on slide, 38 (CNC). Ontario, Carden Alvar, Cameron Ranch, 44.637 -79.058, J. Cossey leg. 6.10.2011, 1 female, BOLD, BIOUG01756 - G05 (CBG). Ontario, Rouge National Urban Park, Toronto Zoo, 43.8223 -79.1897, 125 masl, K. Kerr & A. Sritharan leg. 1.VII.2014, 1 female, BOLD, BIOUG20540 - A09; 1 male, BOLD, BIOUG20540 - B02 (CBG). Ontario, Cambridge, Indian Woods, 43.3736 -80.3652, 304 masl, BIO Collections Staff leg. 14. V.2015, 1 female, BOLD, BIOUG 22361 - C03 (CBG). Ontario, Charleston Lake Provincial Park, Hemlock Ridge Trail, 44.5054 -76.0275, 108 masl, BIObus leg. 22. VI.2015, 1 III instar larva, BOLD, BIOUG23326 - C12 (CBG). Ontario, Owen Sound, Bayview Escarpment Provincial Park, 44.6337 -80.6983, 319 masl, CBG Collections Staff leg. 10.VII.2014, 1 male, BOLD, BIOUG33751 - B08; 1 male, BOLD, BIOUG33752 - A07 (CBG). Ontario, Tiny, Awenda Provincial Park, 44.8253 -79.9846, 231 masl, CBG Collections Staff leg. 8.VIII.2014, 1 female, BOLD, BIOUG34258 - F03 (CBG). Ontario, Morpeth, Rondeau Provincial Park, 42.3021 -81.8531, 239 masl, CBG Collections Staff leg. 13. V.2014, 1 male, BOLD, BIOUG34670 - D06; 1 male, BOLD, BIOUG34670 - G06; 1 female, BOLD, BIOUG34670 - H02 (CBG). Ontario, Perth, Murphy`s Point Provincial Park, 44.7812 -76.2336, 143 masl, CBG Collections Staff leg. 23. V.2014, 1 female, BOLD, BIOUG34683 - F08; 1 male, BOLD, BIOUG34694 - D02 (CBG). Ontario, Warsaw, Ferris Provincial Park, 44.2829 -77.7963, 131 masl, CBG Collections Staff leg. 12.IX.2014, 1 female, BOLD, BIOUG 34924 - D04 (CBG). New Brunswick, Springfield, Belleisle Regional High School, 45.669 -65.836, 13 masl, S. Lawson leg. 2.X.2015, 1 male, BOLD, BIOUG25544 - C01; 1 female, BOLD, BIOUG25544 - F11 (CBG). Prince Edward Island, Prince Edward Island National Park, 46.4123 -63.085, 6 masl, P. Ayles leg. 26. VI.2013, 1 male, BOLD, BIOUG10366 - H09; 1 female, BOLD, BIOUG10368 - A04; 10.VII.2013, 1 male, BOLD, BIOUG10435 - C08; 5. VI.2013, 1 male, BOLD, BIOUG10660 - D02; 1 male, BOLD, BIOUG10787 - B11 (CBG). British Columbia, Kootenay National Park, Kootenay Pond, 50.892 -116.041, 1188 masl, BIObus 2014 leg. 14.VIII.2014, 1 female pupa, BOLD, BIOUG22761 - H04 (CBG). NWT, Nahanni National Park Reserve, Nailicho (Virginia Falls), 61.606 -125.758, 578 masl, Parks Canada leg. 27. VI.2014, 1 male, BOLD, BIOUG16983 - G08; 12.VII.2014, 1 female, BOLD, BIOUG17213 - H03 (CBG). NWT, Reindeer Depot, Mackenzie delta, 68°42’N 134°07’W, J. R. Vockeroth leg. 5.VII.1948, 7 males (5 pinned, 2 on slide) 1 female on slide; 15.VII.1948, 2 females pinned; 1.VIII.1948, 1 female pinned (CNC). USA. Alaska, Big Delta, Boleo Lk., 64.14 -145.81, W. R. M. Mason leg. 22. V.1955, 1 male on slide, 32 (CNC). Alaska, pond near Kuzitrin River, 65.396 -164.496, D. J. Taylor leg. 31.VII.2013, 6 larvae in EtOH, Kuzitrin 4 (UBB). Alaska, pond near Nome Airport, 64.514 -165.422, D. J. Taylor leg. 3.VIII.2011, 4 larvae in EtOH, Nome 1 2011 (UBB). North Carolina, Highlands, 35°03ʹN 83°12ʹW, 3800’, J. R. Vockeroth leg. 24. VI.1957, 4 males (1 on slide, 3 pinned, 1 hypopygium in microvial); 8. VI.1957, 1 female pinned; 10. VI.1957, 1 female pinned; 21. VI.1957, 1 male pinned (CNC). Redescription. Adult male. Head light to dark brown, bearing pale setae. Non-setose area of occiput brownish. Clypeus bearing dark setae. Penultimate flagellomere 285 (271–313), apical flagellomere 204 (163–228), penultimate/apical 1.42 (1.21–1.78, n=6). Lengths of palpal segments 2–5: 105 (89–113), 208 (177–234), 181 (172–194), 336 (267–393, n=6). Thorax. Scutellum and mediotergite orange–dark brown (Fig. 11). Coloration of pleuron composed of pale and dark areas: most of the katepisternum darkened; antepronotal lobe, postpronotum, anepimeron, part of metanepisternum and part of anepisternum slightly darkened, halteres whitish. Thoracic setae (n=6): antepronotal lobe 33 (27–45), postpronotal 4 (3–6), proepisternal 5 (4–7), katepisternal 6 (6–7), anepisternal 16 (12–19), anepimeron 8 (7–9), supra–alar 2 (1–2, n=3). Legs pale yellow–light brown. Foreleg, lengths of fe, ti and ta1–ta5 (n=6, except t4 and t5 n=5): fe 1708 (1667–1750), ti 1773 1722–1859), t1 825 (773–864), t2 499 (455–534), t3 397 (344–432), t4 271 (230–291), t5 196 (177–221). Midleg, lengths of fe, ti and ta1–ta5 (n=6): fe 1467 (1360–1556), ti 1413 (1314–1472), t1 689 (619–727), t2 398 (348–443), t3 318 (261–364), t4 218 (188–235), t5 179 (160–193). Hind leg, lengths of fe, ti and ta1–ta5 (n=6, except fe and t4 n=5): fe 1839 (1723–1945), ti 1761 (1639–1933), t1 1012 (869–1114), t2 581 (454–643), t3 404 (318–443), t4 248 (209–271), t5 185 (171–197). Wing (n=7) : Published as part of Salmela, Jukka, Härmä, Oskari & Taylor, Derek J., 2021, Chaoborus flavicans Meigen (Diptera, Chaoboridae) is a complex of lake and pond dwelling species: a revision, pp. 151-196 in Zootaxa 4927 (2) on pages 169-177, DOI: 10.11646/zootaxa.4927.2.1, http://zenodo.org/record/4536900 : {"references": ["Johannsen, O. A. (1903) Aquatic nematocerous Diptera. 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