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Ursus arctos Brown bear Large carnivores Artificial feeding Cantabrian mountains Feeding sites Food subsidies Food supplementation Movement patterns Supplementary feeding Supplementary food Canis latrans Canis lupus Climate change Habitat fragmentation Habitat use Human dominated landscapes Human-dominated landscapes Human-modified landscapes Human‐modified landscapes Puma concolor Ursus arctos horribilis Ursus maritimus VDP::Zoologiske og botaniske fag: 480 VDP::Zoology and botany: 480 Zoology citizen science genetic sampling grey wolf imperfect detection