Athetina

• Main Authors: Leschen, Richard A. B., Newton, Alfred F.
• Format: Text
• Language: unknown
• Published: Zenodo 2015
• Subjects: Biodiversity; Taxonomy; Animalia; Arthropoda; Insecta; Coleoptera; Staphylinidae; Newfoundland; Canada; New Zealand; Tristan; Baldwin; Fernandez; Chandler; Christchurch; Heer; Holme; Pawson; Kelsey; Siberia
• Online Access: https://dx.doi.org/10.5281/zenodo.6116228; https://zenodo.org/record/6116228

ATHETINA Casey 1910 Acrotona Thomson 1859 Acrotona Thomson 1859: 38. Type species: Aleochara aterrima Gravenhorst, 1802, by original designation; ICZN (1961), Opinion 600. Notes: There are 347 valid species of Acrotona distributed worldwide, with two species apparently adventive in New Zealand. Acrotona aterrima (Gravenhorst 1802) Aleochara aterrima Gravenhorst 1802: 83. Type locality: [Brunswick, Germany]. Type depository: ZMHB? #? ST. Note: New Zealand, Hammond ms in NZAC (identification with "?"). Distribution: Europe, Algeria, Russia (European, Siberia), Georgia, Kazakhstan, Uzbekistan, Korea; New Zealand (intro). Acrotona lugensides Newton 2015 Acrotona lugensides Newton 2015: 9 (replacement name for Homalota lugens Motschulsky 1858: 257). Note: as valid species (as lugens ), not synonym of fungi , Smetana (2004); as adventive in New Zealand (as lugens ), Maddison (2010). Distribution: Indies orientales; intro New Zealand (NZ (intro)). = Homalota lugens Motschulsky 1858: 257 (preoccupied, not Homalota lugens Kiesenwetter 1844 (JPH; JSH in Acrotona )). Type locality: Indes orientales. Type depository: ZMUM? #? ST. Amischa Thomson 1858 Amischa Thomson 1858: 33. Type species: Aleochara analis Gravenhorst, 1802, by monotypy. Notes: There are 60 valid species of Amischa worldwide, with three adventive in New Zealand. Amischa analis (Gravenhorst 1802) Aleochara analis Gravenhorst 1802: 76. Type locality: Brunsuiga [Brunswick, Germany]. Type depository: ZMHB? #? ST. Note: New Zealand, Kuschel (1990), Emberson (1998); redescription, Klimaszewski et al. (2011). Distribution: Europe, Russia (European to Far East), Georgia, Turkey, Iran, Algeria, Tunisia, Azores, Madeira, Kazakhstan, Uzbekistan; intro Canada, USA, Australia, New Zealand, Chatham Is. (AK; MC,NO,SO, CH (intro)). Amischa decipiens (Sharp 1869) Homalota decipiens Sharp 1869: 179. Type locality: England: Chatham; Weymouth; Southend; Hammersmith; Charlton. Type depository: BMNH? #? ST. Note: New Zealand, Kuschel 1990, Sivasubramaniam et al. (1997). Distribution: Europe, Turkey, Tunisia, Canary Is., Madeira; intro. New Zealand (AK,WN; MC (intro)). Amischa nigrofusca (Stephens 1832) Aleochara nigrofusca Stephens 1832: 129 (attributed to Kirby MSS). Type locality: Holme-juxta-mare, in Norfolk [England]. Type depository: BMNH? #? ST. Note: New Zealand, Sivasubramaniam et al. (1997). Distribution: Europe, Russia (n European), Turkey, Algeria, Tunisia, Madeira, Kazakhstan; intro. New Zealand (MC (intro)). Atheta Thomson 1858 Atheta Thomson 1858: 36 (without species). Type species: Aleochara graminicola Gravenhorst, 1806, designated by Thomson (1859: 39); ICZN (1961), Opinion 600. subgenus Alaobia Thomson 1858: 36. Type species: Homalota ochracea Erichson, 1837 (= Atheta scapularis (Sahlberg, C.R., 1831)), by monotypy. subgenus Microdota Mulsant & Rey 1873 a: 160 (without description). Type species: Microdota sericea Mulsant & Rey, 1873 (= Atheta amicula (Stephens, 1832)), designated by Blackwelder (1952: 244). subgenus Ceritaxa Mulsant & Rey 1873 a: 164 (without description). Type species: Ceritaxa spissata Mulsant & Rey, 1873 (= Atheta dilaticornis (Kraatz, 1856)), by monotypy. subgenus Datomicra Mulsant & Rey 1873 b: 355. Type species: Homalota celata Erichson, 1837, designated by Fenyes (1918: 22). subgenus Oreostiba Ganglbauer 1895: 219. Type species: Homalota tibialis Heer, 1839, designated by Fenyes (1918: 24). subgenus Dimetrotina Casey 1911: 143. Type species: Dimetrota vaniuscula Casey, 1911, by monotypy. Notes: There are about 1700 valid species of Atheta worldwide, placed in nearly 90 subgenera, but some of these subgenera are considered separate genera by some authors while other athetine genera (e.g., Acrotona and Philhygra ) are sometimes treated as subgenera of Atheta , so these numbers and the generic/subgeneric concepts are variable. Currently 10 species have been reported from New Zealand, five of them adventive. Atheta (Microdota) amicula (Stephens 1832) Aleochara amicula Stephens 1832: 132 (attributed to Kirby MSS). Type locality: Suffolk [England]. Type depository: BMNH? #? ST. Note: New Zealand, Kuschel (1990), Sivasubramaniam et al. (1997); redescription, Klimaszewski et al. (2011). Distribution: Europe, northern Africa, Canary Is., Madeira, Azores, Russia (European, Siberia), Turkey, Israel, Korea, China; intro Canada, USA, Mexico, Chile, New Zealand, Campbell I. (AK,TO; MC,CA (intro)). Remarks. Type material of Atheta (Atheta) cottieri , A. (Acrotona) muggeridgei and A. (Acrotona) zealandica (synonym of A. pasadenae ) are externally similar and share the same locality labels. Specimens from this series contained in the NZAC were placed under the name A. amicula by Willy Kuschel. Atheta (Atheta) brouni Bernhauer & Scheerpeltz 1926 Atheta (Atheta) Brouni Bernhauer & Scheerpeltz 1926: 639 (new name for Gyrophaena atriceps Broun 1880: 89). Note: AK (as atriceps ), Kuschel (1990); MC (as atriceps ), Bowie et al. (2003). Distribution: New Zealand (AK,ND; MC). = Gyrophaena atriceps Broun 1880: 89 (preoccupied, not Stephens 1832 (JSH in Atheta ); not now congeneric but replaced before 1961 and replacement name in use). Type locality: vicinity of Whangarei Harbour. Type depository: BMNH LT ♂ (des. Klimaszewski et al. 2003: 125) Atheta (Atheta) cottieri Cameron 1945 Atheta (Atheta) cottieri Cameron 1945: 166. Type locality: New Zealand: Palmerston. Type depository: BMNH HT. Note: Kuschel (1990), as intro (origin unknown). Distribution: New Zealand (WI). Remarks. Cameron based his description on a single specimen that we have confirmed as the Holotype. See comment under A. amicula . Type material examined. Holotype (BMNH): card-mounted, “ Type [round label with red border] // from dry rot lesions on swede Palmerston North New Zealand. W. Cottier. Feb 1930 [in Cameron’s hand] // Brit. Mus. 1946 - 60 [in Cameron’s hand] // Atheta (s. str) cotteri TYPE Cam. [in Cameron’s hand, “ type ” written in red ink] // cotteri Cam. i.l. B.M. N. Zeal.”. Atheta kingorum Klimaszewski & Marris 2003 Atheta kingorum Klimaszewski & Marris 2003: 124. Type locality: Three Kings Is. N[ew] Z[ealand]: Great I., Castaway Valley. Type depository: NZAC HT ♂, 17 PT. Distribution: New Zealand (TH (Great I.)). Atheta (Oreostiba) maruiana Cameron 1950 Atheta (Oreostiba) maruiana Cameron 1950: 24. Type locality: Westland: Maruia Springs, South Island. Type depository: BMNH LT; NZAC 6 PLT (designated below). Distribution: New Zealand (BR). Remarks. Cameron did not indicate the number of specimens he examined to name this species. One specimen was located in the BMNH and several in the NZAC, two of which were placed under the combination “ Amriathaea antipoda (Bernhauer 1943)” by Kuschel. In order to stabilize this name a lectotype and six paralectotypes are here designated for the specimens of Atheta (Oreostiba) maruiana . Type material examined. Lectotype (BMNH): card-mounted, “SYN-TYPE [round label with powder blue border] // Westland N. Zealand Maruia Springs IX. 1948 [in Cameron’s hand] // A (Oreostiba) maruiana Cam. TYPE [in Cameron’s hand, “ type ” written in red ink] // Atheta maruana Cam [in Bacchus’ hand] M. E. Bacchus det 1971 SYNTYPE // M. Cameron. Bequest. B.M. 1955 - 147.”. Paralectotypes (NZAC): 4 card-mounted (2 on one card, another card that has one specimen but with 2 specimens missing), “Maruia Springs Westland South Is. N. Z.[in Brookes’ hand] // coll. Brookes Sept. 1948. [in Brookes’ hand] // taken from leaf-mould [in Brookes’ hand] // Atheta. (Oreostiba) maruiana Cameron. [in Brookes’ hand] // Paratype [in Cameron’s? hand, pink underline] // A. E. Brookes Collection // Paratype [dark blue label]; 2 (card-mounted on same card), “Maruia Springs Westland South Is. N. Z.[in Brookes’ hand] // coll. Brookes Sept. 1948. [in Brookes’ hand] // taken from leaf-mould [in Brookes’ hand] // A (Oreostiba) maruiana Cameron. Type [in Cameron’s hand. “ type ” in red ink] // A. E. Brookes Collection // NZAC specimen database record 1007110 // NZ Arthropod Collection Private Bag 92170 Auckland New Zealand NZAC 04065749 [barcode label]”. Atheta (Datomicra) muggeridgei Bernhauer 1943 Atheta (Datomicra) Muggeridgei Bernhauer 1943: 176. Type locality: Palmerston. Type depository: FMNH LT, 1 PLT; BMNH 2 PLT (designated below). Distribution: New Zealand (WI). Remarks. Bernhauer did not indicate the number of specimens he examined. Four specimens labelled as types were located in the BMNH and the FMNH, and a fifth specimen in the BMNH only had an identification label handwritten by Bernhauer that was not considered a syntype. In order to stabilize this name, a lectotype (FMNH) and three paralectotypes (BMNH, FMNH) are here designated from this material of Atheta (Datomicra) muggeridgei . See comment under A. amicula . Type material examined. Lectotype (FMNH): card-mounted, “dry rot lesions on swede. Palmerston Nth. W. Cottier 25 / 3 / 30 [in Cameron’s hand] // New Zealand Palmerston Nth 25 March 1930 [in Bernhauer’s hand] // Muggeridgei Bernh. Typ. Microdota [in Bernhauer’s hand] // Muggeridgei Brnh Typus Datomicra [in Bernhauer’s hand; red label] // Chicago NHMus M.Bernhauer Collection // SYNTYPE teste D. J. Clarke 2014 GDI Imaging Project [violet label] // Photographed Kelsey Keaton 2014 Emu Catalog [blue label] // FMNHINS 2819065 FIELD MUSEUM [barcode label]”. Paralectotypes: 1 (FMNH): card-mounted, “dry rot lesions on swede. Palmerston Nth. W. Cottier 25 / 3 / 30 [in Cameron’s hand] // Atheta Muggeridgei Brnh. n. sp. [in Bernhauer’s hand] // Muggeridgei Brnh. Cotypus Datomicra [in Bernhauer’s hand; red label] // Chicago NHMus M.Bernhauer Collection // SYNTYPE teste D. J. Clarke 2014 GDI Imaging Project [violet label]”; 1 (BMNH): pointed, “ Type [round label with red border] // dry rot lesion on swede. Palmerston Nth W. Cottier 25 / 1 / 30 [in Cameron’s hand] // Atheta Muggeridgei Brnh. n. sp. [in Bernhauer’s hand]”; 1 (BMNH): pointed, “Co-type [round label with yellow border] / / dry rot lesion on swede. Palmerston Nrth W. Cottier 25 / 1 / 30 [in Cameron’s hand] // Pres. By Imp. Inst. Ent. Brit. Mus. 1931 - 288.”. Atheta (Datomicra) nigra (Kraatz 1856) Homalota nigra Kraatz 1856: 287. Type locality: Wohl über ganz Deutschland. Type depository: SDEI? #? ST. Note: New Zealand, Sivasubramaniam et al. (1997). Distribution: Europe, Russia (European), Turkey, Azores, Canary Is., Korea, China (Yunnan); intro. New Zealand (MC (intro)); intro. Canada (Klimaszewski et al. 2015). Atheta (Dimetrotina) pasadenae Bernhauer 1906 Atheta (Acrotona) pasadenae Bernhauer 1906: 338. Type locality: [U.S.A.]: California: Pasadena. Type depository: FMNH LT ♂, 2 PLT (des. Gusarov (2015)). Note: Species previously reported from New Zealand as Atheta zealandica or other synonyms below: Kuschel (1990) as intro.; MC, Bowie et al. (2003); Palaearctic distribution, Assing & Schulke (2006: 112). Distribution: Inaccessible, Tristan da Cunha & Nightingale Is.,? South Africa; intro? USA (CA,NV), Chile, Juan Fernandez Is., Hawaii (Kauai), France, n Africa, Azores, Madeira, Canary Is., New Zealand (AK,MC; WI (intro)). = Atheta (Acrotona) zealandica Cameron 1945: 167. Type locality: New Zealand: Palmerston. Type depository: BMNH LT ♂, 1 PLT (des. Leschen in Gusarov (2015)). Note: synonym of A. pasadenae , Gusarov (2015). = Atheta (Atheta) pseudocoriaria Bernhauer 1943: 175 (preoccupied, not Atheta pseudocoriaria Cameron 1939 (JPH)). Type locality: Nord-Insel: Palmerston; Auckland. Type depository: BMNH LT ♂, 4 PLT (des. Leschen in Gusarov (2015)); FMNH 6 PLT; NZAC 6 PLT. Note: implied synonym of zealandica, Cameron (1945), Kuschel (1990: 46). = Atheta (Atheta) aucklandensis Pace 1987: 436 (new name for Atheta pseudocoriaria Bernhauer 1943: 175). Remarks. The species concept, synonymy, subgeneric assignment and world distribution of this species has been revised by Gusarov (2015), who discussed additional synonyms not reported from New Zealand. Below we have indicated 12 additional paralectotypes in FMNH and NZAC not examined or listed by Gusarov (2015). Type material examined. Atheta (Atheta) pseudocoriaria Bernhauer. Paralectotypes: 6 pinned with minutens (NZAC, one specimen with an identification label, “ Atheta pseudocoriaria CoType. Bern” in unknown hand), “Co- type [round label with yellow border] // X dry-rot swede Mr. Gibbs M. C. area Palm. Nth. N. Z. W. Cottier 25 / 6 / 31 [in unknown hand] // A [handwritten]”; 1 card-mounted (FMNH), “X dry-rot swede Mr. Gibbs M. C. area Palm. Nth. N. Z. W. Cottier 25 / 6 / 31 [in Cameron’s hand?] // pseudocoriaria Bernh. Typus [in Bernhauer’s hand, red label] // Chicago NHMus M.Bernhauer Collection // SYNTYPE teste D. J. Clarke 2014 GDI Imaging Project [violet label] // Photographed Kelsey Keaton 2014 Emu Catalog [blue label] // FMNHINS 2819083 FIELD MUSEUM [barcode label]”; 2 (FMNH) card-mounted [1 missing abdomen]: “X dry-rot swede Mr. Gibbs M. C. area Palm. Nth. N. Z. W. Cottier 25 / 6 / 31 [in Cameron’s hand?] // A [handwritten] // pseudocoriaria Bernh. Cotypus Atheta s. str. [in Bernhauer’s hand, red label] // Chicago NHMus M.Bernhauer Collection // SYNTYPE teste D. J. Clarke 2014 GDI Imaging Project [violet label]”; 1 (FMNH) card-mounted: “dry-rot lesions on swede // Palmerston Nth. W. Cottier 25 / 3 / 30 [in Cameron’s hand?] // Atheta zealandica Brnh. n. sp. [in Bernhauer’s hand] // pseudocoriaria Bernh. Cotypus Atheta s. st. [in Bernhauer’s hand, red label] // Chicago NHMus M.Bernhauer Collection // SYNTYPE teste D. J. Clarke 2014 GDI Imaging Project [violet label]”; 1 (FMNH) card-mounted: “Aucklnd N.Z [handwritten, oval red label] // Auckland. [red line] New Zealand. // Sharp Coll 1905 - 313 // Atheta pseudocoriaria Brnh. [in Bernhauer’s hand] Bernhauer det. // BRIT. MUS. DON. ARROW // pseudocoriaria Bernh. Cotypus Atheta s. st. [in Bernhauer’s hand, red label] // Chicago NHMus M.Bernhauer Collection // SYNTYPE teste D. J. Clarke 2014 GDI Imaging Project [violet label]”; 1 (FMNH) card-mounted [in 3 pieces, antennae and legs missing]: “Auckland Lawson [handwritten] // Auckland, New Zealand. // 19. [handwritten] // Sharp Coll 1905 - 313 // BRIT. MUS. DON. ARROW // pseudocoriaria Bernh. Cotypus [in Bernhauer’s hand, red label] // Chicago NHMus M.Bernhauer Collection // SYNTYPE teste D. J. Clarke 2014 GDI Imaging Project [violet label]”. Atheta (Ceritaxa) splendidicollis Bernhauer 1943 Atheta (Ceritaxa) splendidicollis Bernhauer 1943: 175. Type locality: Nord-Insel. Type depository: BMNH LT; FMNH 1 PLT (designated below). Note: as implied synonym of Atheta (now Leptostiba ) politula , Kuschel (1990). Distribution: New Zealand (NO). Remarks. Bernhauer did not indicate the number of specimens he examined. Two specimens were hand-labelled as types in the BMNH and FMNH. In order to stabilize this name a lectotype (BMNH) and one paralectotype (FMNH) are here designated from material of Atheta (Ceritaxa) splendidicollis . Type material examined. Lectotype (BMNH): card-mounted, “ Type [round label with red border] // North Island, N. Zealand. R. N. Parker. // G.C. Champion Coll. B.M. 1927 - 409. // Atheta splendi-collis [ sic ] Brnh. Typus [in Bernhauer’s hand] M. Bernhauer det. //”. Paralectotype (FMNH): card-mounted, “North Island, N. Zealand. R. N. Parker. // G.C. Champion Coll. B.M. 1927 - 409. // Atheta splendidicollis Brh. Typ. [in Bernhauer’s hand] Bernhauer det. // BRIT. MUS. DON. ARROW // splendidicollis Brnh. Typus Ceritaxa. [in Bernhauer’s hand, red label] // Chicago NHMus M.Bernhauer Collection // SYNTYPE teste D. J. Clarke 2014 GDI Imaging Project [violet label] / / Photographed Kelsey Keaton 2014 Emu Catalog [blue label] // FMNHINS 2819077 FIELD MUSEUM [barcode label]”. Atheta (Alaobia) trinotata (Kraatz 1856) Homalota trinotata Kraatz 1856: 272 (should be protected over 2 senior names, teste Smetana (2004: 33)). Type locality: [no locality cited; Deutschland implied]. Type depository: SDEI? #? ST. Note: New Zealand, Hammond ms. Distribution: Europe, Russia (European), Georgia, Ukraine, Turkey, Iran, Cyprus, Algeria, Morocco, Tunisia, Canary Is., Madeira; intro. New Zealand (DN (intro)). Brundinia Tottenham 1949 Brundinia Tottenham 1949: 78 (new name for Metaxya Mulsant & Rey). Type species: Metaxya apricans Mulsant & Rey, 1873 (= Brundinia meridionalis (Mulsant & Rey, 1853)), by autotypy. = Metaxya Mulsant & Rey 1873 a: 181 (preoccupied, not Walker 1856; without description). Type species: Metaxya apricans Mulsant & Rey, 1873 (= Brundinia meridionalis (Mulsant & Rey, 1853)), designated by Blackwelder (1952: 240). Notes: Brundinia includes 18 valid species scattered worldwide, with one endemic to New Zealand. Brundinia semipallidula (Bernhauer 1943) Atheta (Metaxya) semipallidula Bernhauer 1943: 174. Type locality: Wedderburn. Type depository: FMNH LT; BMNH 2 PLT (designated below). Distribution: New Zealand (CO). Remarks. Bernhauer did not specify the exact number of specimens he examined for naming Atheta (Metaxya) semipallidula . There were two specimens mounted on the same card with a label hand-written “ typus ” in the BMNH and a single specimen in the FMNH. In order to stabilize this name a lectotype (FMNH) and 2 paralectotypes are here designated from material labelled as types of Atheta (Metaxya) semipallidula . Type material examined. Lectotype (FMNH): card-mounted, “ New Zealand S : Published as part of Leschen, Richard A. B. & Newton, Alfred F., 2015, Checklist and type designations of New Zealand Aleocharinae (Coleoptera: Staphylinidae), pp. 301-353 in Zootaxa 4028 (3) on pages 305-312, DOI: 10.11646/zootaxa.4028.3.1, http://zenodo.org/record/237988 : {"references": ["Casey, T. L. (1910) New species of the staphylinid tribe Myrmedoniini. In: Memoirs on the Coleoptera. Vol. 1. New Era Printing Co., Lancaster, Pennsylvania, pp. 1 - 183.", "Thomson, C. G. (1859) Skandinaviens Coleoptera, synoptiskt bearbetade. I. Berlingska Boktryckeriet, Lund, 290 pp.", "Gravenhorst, J. L. C. 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