Ariadna boesenbergi Keyserling 1877
Ariadna boesenbergi Keyserling, 1877 Figures 22‒23, 70 Ariadne bösenbergii Keyserling, 1877: 223, pl. 7, fig. 7 (♀ lectotype and ♂ lectoparatype from Montevideo [34°53'01"S; 56°10'55"W], Uruguay, Bösenberg leg., deposited in ZMH 58, designated by Beatty (1970)...
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Online Access: | https://dx.doi.org/10.5281/zenodo.5987870 https://zenodo.org/record/5987870 |
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Open Polar |
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Biodiversity Taxonomy Animalia Arthropoda Arachnida Araneae Segestriidae Ariadna Ariadna boesenbergi |
spellingShingle |
Biodiversity Taxonomy Animalia Arthropoda Arachnida Araneae Segestriidae Ariadna Ariadna boesenbergi Giroti, André Marsola Brescovit, Antonio Domingos Ariadna boesenbergi Keyserling 1877 |
topic_facet |
Biodiversity Taxonomy Animalia Arthropoda Arachnida Araneae Segestriidae Ariadna Ariadna boesenbergi |
description |
Ariadna boesenbergi Keyserling, 1877 Figures 22‒23, 70 Ariadne bösenbergii Keyserling, 1877: 223, pl. 7, fig. 7 (♀ lectotype and ♂ lectoparatype from Montevideo [34°53'01"S; 56°10'55"W], Uruguay, Bösenberg leg., deposited in ZMH 58, designated by Beatty (1970), examined; ♂ type from Uruguay, deposited in NHM 90.7.1.475-6-7, examined). Ariadna bösenbergi — Petrunkevitch (1911: 130).— Mello-Leitão (1943: 233). Ariadne bösenbergi — Gerhardt (1921: 92). Ariadna mollis— Mello-Leitão (1933: 12, Syn. with A. boesenbergi 1947: 234).— Bonnet (1955: 730, 735). Ariadna boesenbergi— Beatty (1970: 476, remove the syn. with A. mollis , descr. ♀).— Prandi (1990: 3).— Grismado (2008: 357, figs 8N, O, 10E, H).— Ramírez (2014: 211, figs 139A, 141D, 167A).— Grismado & Izquierdo (2014: 154, figs 8, 10a‒b, 12).— World Spider Catalog (2018). Additional material examined. BRAZIL: Rio Grande do Sul State: Itaqui /Maçambará, Reserva Biológica São Donato [29°02'46,7"S; 56°04'48,7"W], 1♂, 23.I.2012, M. Machado leg. (IBSP 166778); Quaraí [30°23'17"S; 56°29'56"W], Cerro do Jarau, 1♀, 23.II.1975, A.A. Lise leg. (MCN 02508); Rio Grande [32°02'06"S; 52°05'55"W], 1♀ 4imm. (NHM 89.2.17; 90.7.1.477); ARGENTINA: Tucumán Province: Arroyo India Muerta, Road to Ticucho [26°31'S; 65°15'W], 1♀, 25.VI.1995, M. Ramírez & P. Goloboff leg. (MACN-Ar 10161); La Rioja Province: Chilecito [29°10'S; 67°30'W], 1♀, 27‒30.I.1956, M.E. Galiano leg. (MACN-Ar 10163); Entre Ríos Province: Concordia [31°24'S; 58°02'W], under bark of Melia azedarach , 1♀, X.1992, C. Grismado leg. (MACN-Ar 10175); Córdoba Province: Córdoba [31°25'S; 64°11'W], Bosque Alegre, 3♀ 2imm., IX.1981, P. Goloboff leg. (MACN-Ar 10166; 10167; 10168); Nono, 880m (31°47'44.3"S; 64°00'6,26"W), 2♀ 3imm., 3‒9.II.2008, M. Ramírez leg. (MACN-Ar 13504; 13505); Leones [32°42'21"S; 62°06'30"W], 1♀, 13.X.1946, W. Patridge leg. (MACN-Ar 10164); 1♀, 12.X.1946 (MACN-Ar 10234); Santa Fé Province: Santa Fé [31°39'00"S; 60°43'60"W], 2♂, I.1965, M. Aschenbach leg. (MACN-Ar 5853); 1♀, 9.X.1963, Pignalloni leg. (MACN-Ar 10176); San Luís Province: Cortaderas (in wood of " Molles ", Schinus sp., under stones) (32°29'58,9"S; 65°00'01,5"W), 1♀, 10.XI.1982, A. Roig leg. (MACN-Ar 10169); Carolina [32°48'S; 66°06'W], between stones, 3♀, XI.1992, C. Scioscia leg. (MACN-Ar 10171; 10172); Quebrada Merlo, 3♀ 1imm., 12.XI.1982, A. Roig leg. (MACN-Ar 10170); Sierra Central, 3♀ (Ex-MNHNP-AR 9073); Buenos Aires Province: San Pedro [33°40'46"S; 59°40'01"W], in a tube under bark of Eucalyptus, 1♀, 24.XII.1912, A.G. Frers leg. (MACN-Ar 10187); Pergamino [33°44'S; 60°36'W], 2♀, IX.2007, S. Rodriguez Gil leg. (MACN-Ar 35097; 35105); Fontezuela [33°55'S; 60°28'W], 5♀, IX.2007, S. Rodriguez Gil leg. (MACN-Ar 35089; 35094; 35096; 35098; 35100); Ruta Pergamino- Arrecifes, km 210, 5♀, IX.2001, S. Rodriguez Gil leg. (MACN-Ar 28170; 35099; 35102; 35103; 35104); Arrecifes [34°02'S; 60°10'W], Todd, 1♀, IX.2007, S. Rodriguez Gil leg. (MACN-Ar 35106); Zelaya [34°21'38"S; 58°52'36"W], 2♀ 1imm., VIII.1939, H. Hepper leg. (MACN-Ar 10188; 10189; 10231); Tigre [34°25'S; 58°35'W], 1♂, I.1938, J.M. Viana leg. (MACN-Ar 303); Martínez [34°29'S; 58°30'W], 3♂ 4♀ 4imm., V.1976, R. Maynard leg. (MNHNU 1036); Saavedra [34°33'18,30"S; 58°29'32,67"W], 1♀, VIII.1937, Castillo leg. (MACN-Ar 706); Junín [34°35'38"S; 60°56'47"W], 1♀, II.1962, Daguerre leg. (MACN-Ar 10186); Buenos Aires [34°35'59"S; 58°22'55"W], Capital Federal, 7♀, 1967, Bachmann leg. (MACN-Ar 21349); 4♀, VII.1945 (MACN-Ar 1585); 2♂, I.1989, M. Miranda leg. (MACN-Ar 10177; 21308); 3♀ 1imm., VIII.1948, D.Y. Carpintero & O. Carpintero leg. (MACN-Ar 10178); 1♀, VI.1964, A. Martínez leg. (MACN-Ar 10180); 4♀, 1967, Bachmann leg. (MACN-Ar 21309); 1♂, 30.XII.2008, C. Kopuchian leg. (MACN-Ar 21312); 4♀ 1imm., Bery leg. (MNHNP-AR 7045); 1♀ 2imm., 1‒31.XII.1968, Bachmann leg. (MNHNP-AR 14916); Parque Patrícios, 1♀, X.1948 (MACN-Ar 10179); Parque Centenário, under bark of Tipuana tipu , 1♀, X.1992, C. Grismado leg. (MACN-Ar 10182); Vila Lugano, 1♀ 1imm., L.R. Rodriguez leg. (MACN-Ar 10183; 10184); bit to Paladino, 1♂, XII.1965 (MACN-Ar 10185); Arbe frente ao Museo de Ciencias Naturales, 1♀ 4imm., 9.XI.1960 (MACN-Ar 21 360); Jardines del Museo de Ciencias Naturales, 1♀, III.2010, C. Grismado & M. Izquierdo leg. (MACN-Ar 2 9449); Hurlingham, Arroyo Morón [34°36'S; 58°38'W], 2♀ 1imm., XI.1962, E. Massoia leg. (MACN-Ar 10190); Moreno [34°39'04"S; 58°47'24"W], 1♀ 1imm., III.1942, L.R.D. Schiapelli leg. (MACN-Ar 2354); 2♀ 1imm., III.1 947, Exp. R.D. Schiapelli leg. (MACN-Ar 10191); 2♀, X.1947, R.D. Schiapelli leg. (MACN-Ar 21310); Sarandí, Plaza del Viaducto [34°39'45"S; 58°21'54"W], 3♀ 34imm., 30.IX.2007, C. & A. Grismado leg. (MACN-Ar 21311; 21313; 21314); Sarandí [34°41'S; 58°20'W], on wall of bricks, 3♀ 2imm., 27.IV.2002, C., D., & A. Grismado leg. (MACN-Ar 10204); Sarandí, 1♀, 15.VII.2006, C. Grismado & A. Raya leg. (MACN-Ar 11425); Villa Madero [34°42'S; 58°30'W], 5♀, VI.2007, C. Scioscia leg. (MACN-Ar 35090; 35091; 35095; 35101; 35107); 2♀, VIII.2007 (MACN-Ar 35092; 35093); Florencio Varela [34°47'51"S; 58°16'34"W], 1♀, XII.1939, F. Monrós leg. (MACN-Ar 10197); Glew [34°53'17"S; 58°23'03"W], 1♀, 1970, Carpintero leg. (MACN-Ar 10194); 1♀ 1imm., 1979 (MACN-Ar 10196); 1imm., 1972 (MACN-Ar 10195); Tandil [37°19'43"S; 59°08'13"W], La Cascada, 1♀, 16.V.1973, C. Cesari leg. (MACN-Ar 10200); Tandil, 1♀ 3imm., V.1967, E. Maury leg. (MACN-Ar 10199); Sierra de los Padres [37°57'S; 57°46'W], 2♀ 2imm., XI.1962, M.E. Galiano leg. (MACN-Ar 10202); Tornquist, Estancia Fortin Chaco [38°06'S; 62°13'W], 1♂, 1‒31.I.1972, J.A. Obarrio leg. (MNHNP-AR 14915); 15km O de Lobería [38°08'59"S; 58°47'00"W], 1♀, 4.IX.1972 (MACN-Ar 10127); Río Negro Province: General Fernandez Oro [38°57'S; 67°55'W], 1♂, I.1976, Coscaron leg. (AMNH); URUGUAY: Without specific data: 3♀ 1imm. (MACN- Ar 21361); Cerro Largo Department: Ruta 8, Rio Tacuari, 2♀, 15.IV.1965, F. Achaval leg. (MNHNU 1050); Río Negro Department: San Javier [32°41'S; 58°08'W], 4♀, VI.1962, L.C. Zolessi leg. (MNHNU 569); Durazno Department: Durazno, Ruta 5, km 172 [33°24'47"S; 52°30'02"W], 6♀ 4imm., 15.V.2009, M. Simó y col. leg. (MNHNU FCE 3152); Lavalleja Department: Minas [34°22'39"S; 55°14'17"W], 2♀ 1imm., 5.I.1990, L. Prandi leg. (MNHNU 1070); Colonia Department: Colonia del Sacramento [34°38'05"S; 57°50'27"W], 6♂ 9♀ 1imm., 27.VIII.1989, L. Prandi leg. (MNHNU 391); San José Department: Ciudad del Plata, Playa Pascual [34°44'43"S; 56°27'07"W], 1♀ 5imm., 13.I.1990, L. Prandi leg. (MNHNU 540); Canelones Department: Parque Natural INIA Las Brujas [34°38'S; 56°20'W], 4♀, 6.VI.2013, A.D. Brescovit leg. (IBSP 166779); Montevideo Department: Montevideo, 7ª Sec. Judicial [34°53'01"S; 56°10'55"W], 1♂, VI.1964, R. Capocasale leg. (CAS 9039513); Montevideo, Buceo [34°53'59"S; 56°07'58"W], 21♀ 5imm., I.1976, R. Maynard leg. (MNHNU 575; Ex-MNHNU 656); Montevideo, Cordón [34°54'00"S; 56°10'50"W], 2♂, VI.1964, R. Capocasale leg. (MNHNU 1020). Diagnosis. Ariadna boesenbergi is similar to A. crassipalpa , A. mollis , A. calilegua , A. changellkuk, A. levii and A. perkinsi due to the dark abdominal pattern with arrow-shaped marks (Figs 1A‒C, 18A, F, 20A, F, 22A, F, 31A, E, 45A, F, 47A, F, 50A, F), but differ, except A. changellkuk , by the presence of males with metatarsi I straight, without modified macrosetae (Fig. 22C‒E); it differs from A. changellkuk by having a less elongated body (Fig. 22A‒B) and a more elongated mid-portion and embolus (Fig. 23A‒B); females are distinguished by the presence of a unillobulated anterior receptaculum, flattened laterally (Fig. 23C‒D). Description. Male (IBSP 166778). Carapace and chelicerae reddish brown (Fig. 22A). Endites brownish yellow; labium dark brownish yellow. Sternum brownish yellow, with darker margins, marbled with dark pigment, and three pairs of dark marginal areas, the hind one fainted (Fig. 22B). Palps brownish orange (Fig. 23A‒B). Legs brownish yellow, pairs I‒II darker with dark pigment forming diffuse rings in the distal region of metatarsus and tarsus (Fig. 22C‒E). Abdomen withish grey, with a conspicuous dark dorsal pattern comprising arrow-shaped marks (Fig. 22A‒B). Total length 6.64; carapace 3.4 long, 2.16 wide. Chelicerae not projected (Fig. 22A). Palps: bulb elliptical, mid-portion slightly shorter than the embolus; embolus tubular, tapered, and abruptly curved; tibia short and slightly incrassated, almost cylindrical, with the dorso-ventral diameter almost equal to the bulb (Fig. 23A‒B). Legs: robust and coated with setae (Fig. 22C‒E); tarsal brush of putative chemosensory setae present in legs I‒IV, but reduced in legs I, III and IV. Leg formula: I-II-IV-III. Leg measurements: I, femur 3.16, patella 1,.2, tibia 2.6, metatarsus 3.04, tarsus 0.8, total 10.8; II, 2.88, 1.16, 2.6, 2.8, 0.76, 10.2; III, 2.0, 0.96, 1.56, 1.84, 0.64, 7.0; IV, 2.56, 1.24, 2.04, 2.0, 0.6, 8.44. Macrosetae: I, femur D0-0-0-1-1-1/0-2-0, P0-0-0-0-0-2-1-0; tibia P0-1-0-1- 0-1-0, vp1-0-1-0-1-0-1, vr2-0-1-0-1-0-1, R1-1-1-1-1-1-0; metatarsus p0-1-0-0-0-0-0-0, vp0-1-0-0-1-0-0-1, vr0-1- 0-0-1-0-0-1, r0-1-0-0-0-0-0-0 (Fig. 22C‒E); II, femur D0-0-1-1-1-0-2-0, P0-0-0-0-0-1-1-0; tibia P0-1-0-1-0-1-0, vp1-0-1-0-1-0-1, vr2-0-1-0-1-0-1, R1-1-0/1-1-0/1-1-0; metatarsus p0-1-0-0-0-0-0-0, vp0-1-0-0-1-0-0-1, vr0-1-0-0- 1-0-0-1, r0-1-0-1-0-1-0-0; IV, femur D0/1-1-1-0-1-1-0-1-0-0-0; tibia vp0-0/1-0-0-0-0-0, vr0-0-0-0-0-0-0/1; metatarsus vp0-0-0/1-0-0-0-2, vr metatarsal comb with 4 macrosetae. Female (IBSP 166779). Carapace and chelicerae brown, with cephalic region darker (Fig. 22F). Endites brown; labium dark brown. Sternum brownish orange, with darker margins, marbled with dark pigment, and three pairs of dark marginal areas (Fig. 22G). Palps brownish yellow, with tibia and tarsus reddish. Legs brownish yellow, with pairs I‒II darker. Metatarsus and tarsus of legs I‒II darker than the other segments. Dark pigment forming diffuse rings in the distal region of tibia, metatarsus and tarsus, of legs I‒II (Fig. 22I ‒J). Abdomen as in male (Fig. 22F‒G). Total length 9.88; carapace 4.44 long, 2.68 wide. Chelicerae not projected (Fig. 22F). Legs: robust and densely coated with setae, mainly on the ventral region of pairs I‒III; legs I‒II with ventro-prolateral and ventro-retrolateral tibial macrosetae of similar size, and metatarsal macrosetae of different sizes (Fig. 22I ‒J). Leg formula: I-II-IV-III. Leg measurements: I, femur 3.04, patella 1.52, tibia 2.36, metatarsus 2.32, tarsus 0.76, total 10.0; II, 2.76, 1.48, 2.32, 2.16, 0.68, 9.4; III, 2.16, 1.24, 1.48, 1.52, 0.6, 7.0; IV, 2.8, 1.56, 2.16, 1.8, 0.64, 8.96. Macrosetae: I, femur P0-0-0-0-0-1-0; tibia vp1-0-1-0-1-0-1, vr1-0-1-0-1-0-1, R0-1-0-0-0-0-0; metatarsus vp1-1-0/ 1-1-1-1-1-1, vr1-1-1-1-1-1-1 (Fig. 22I ‒J); II, femur D0-0-0-0-0-0/2-0, P0-0-0-0-1-1-0; tibia P0-0-0-0/1-0-0/1-0, vp1-0-1-0-1-0-1, vr1-0-1-0-1-0-1; metatarsus vp1-1-0/1-1-1-1-1-1, vr0/1-1-1-1-1-1-1-1-1; IV, femur D0-1-1-0-0- 0-0; metatarsus vp0-0-0-0-0-0-1, vr metatarsal comb with 4 macrosetae. Genital external plate wide and conspicuous, occupying 1/3 of the epigastric region (Fig. 22G‒H). Internal genitalia: anterior receptaculum with the glandular plate rounded (Fig. 23C‒D). Variation. Male (n=10): Total length 5.12‒7.04; carapace 2.92‒3.52 long, 2.0‒2.48 wide; femur I 2.44‒3.4. Female (n=10): Total length 8.72‒11.64; carapace 3.84‒4.88 long, 2.44‒3.08 wide; femur I 2.64‒3.4. Distribution. South of Brazil, Uruguay, and center and east of Argentina (Fig. 70). : Published as part of Giroti, André Marsola & Brescovit, Antonio Domingos, 2018, The taxonomy of the American Ariadna Audouin (Araneae: Synspermiata: Segestriidae), pp. 1-114 in Zootaxa 4400 (1) on pages 37-41, DOI: 10.11646/zootaxa.4400.1.1, http://zenodo.org/record/1207641 : {"references": ["Keyserling, E. (1877) Amerikanische Spinnenarten aus den Familien der Pholcoidae, Scytodoidae und Dysderoidae. Verhandlungen der Zoologisch-Botanischen Gesellschaft in Wien, 27, 205 - 234.", "Beatty, J. A. (1970) The spider genus Ariadna in the Americas (Araneae, Dysderidae). Bulletin of the Museum of Comparative Zoology, 139 (8), 433 - 518.", "Petrunkevitch, A. (1911) A synonymic index-catalogue of spiders of North, Central and South America with all adjacent islands, Greenland, Bermuda, West Indies, Terra del Fuego, Galapagos, etc. Bulletin of the American Museum of Natural History, 29, 1 - 791. https: // doi. org / 10.5962 / bhl. title. 23819", "Mello-Leitao, C. F. de (1943) Catalogo das aranhas do Rio Grande do Sul. Arquivos do Museu Nacional do Rio de Janeiro, 37, 147 - 245.", "Gerhardt, U. (1921) Vergleichende Studien uber die Morphologie des mannlichen Tasters und die Biologie der Kopulation der Spinnen. Archiv fur Naturgeschichte, 87, 78 - 247.", "Mello-Leitao, C. F. de (1933) Catalogo das aranhas argentinas. Archivos da Escola Superior de Agricultura e Medicina Veterinaria, Rio de Janeiro, 10, 3 - 63.", "Bonnet, P. (1955) Bibliographia araneorum. 2 (1). Les Artisans de L'Imprimerie Douladoure, Toulouse, 919 pp.", "Prandi, L. (1990) El genero Ariadna (Araneae, Segestriidae) en la Republica Oriental del Uruguay. Arachnologia, 5 (Supplement), 1 - 10.", "Grismado, C. J. (2008) A taxonomic revision of the spider genus Ariadna Audouin, 1826 in Argentina and Chile, with the description of five new species (Arachnida, Araneae, Segestriidae). Zoosystema, 30 (2), 333 - 360.", "Lipke, E., Ramirez, M. J., Michalik, P. (2014) Ultrastructure of spermatozoa of Orsolobidae (Haplogynae, Araneae) with implications on the evolution of sperm transfer forms in Dysderoidea. Journal of Morphology, 275, 1238 - 1257. https: // doi. org / 10.1002 / jmor. 20298", "Grismado, C. J. & Izquierdo, M. A. (2014) Dysderoidea. In: Roig-Junent, S., Claps, L. E. & Morrone, J. J. (Eds.), Biodiversidad de Artropodos Argentinos. Vol. 3. Sociedad Entomologica Argentina, La Plata, pp. 151 - 166.", "World Spider Catalog (2018) World Spider Catalog. Natural History Museum Bern, version 19.0. Avaliable from: http: // wsc. nmbe. ch / (accessed on 16 January 2018)"]} |
format |
Text |
author |
Giroti, André Marsola Brescovit, Antonio Domingos |
author_facet |
Giroti, André Marsola Brescovit, Antonio Domingos |
author_sort |
Giroti, André Marsola |
title |
Ariadna boesenbergi Keyserling 1877 |
title_short |
Ariadna boesenbergi Keyserling 1877 |
title_full |
Ariadna boesenbergi Keyserling 1877 |
title_fullStr |
Ariadna boesenbergi Keyserling 1877 |
title_full_unstemmed |
Ariadna boesenbergi Keyserling 1877 |
title_sort |
ariadna boesenbergi keyserling 1877 |
publisher |
Zenodo |
publishDate |
2018 |
url |
https://dx.doi.org/10.5281/zenodo.5987870 https://zenodo.org/record/5987870 |
long_lat |
ENVELOPE(-67.700,-67.700,-67.183,-67.183) ENVELOPE(-62.300,-62.300,-64.083,-64.083) ENVELOPE(-62.183,-62.183,-64.650,-64.650) ENVELOPE(-85.933,-85.933,-78.050,-78.050) ENVELOPE(-60.583,-60.583,-63.033,-63.033) ENVELOPE(-63.200,-63.200,-64.683,-64.683) ENVELOPE(-62.233,-62.233,-63.250,-63.250) ENVELOPE(-56.720,-56.720,-63.529,-63.529) ENVELOPE(18.412,18.412,69.835,69.835) ENVELOPE(-67.050,-67.050,-68.217,-68.217) ENVELOPE(-56.683,-56.683,-63.583,-63.583) ENVELOPE(-57.931,-57.931,-63.317,-63.317) ENVELOPE(-56.950,-56.950,-64.333,-64.333) ENVELOPE(-60.583,-60.583,-62.933,-62.933) ENVELOPE(-66.333,-66.333,-66.433,-66.433) ENVELOPE(-41.417,-41.417,-82.250,-82.250) ENVELOPE(-63.417,-63.417,-65.117,-65.117) ENVELOPE(-60.800,-60.800,-62.450,-62.450) ENVELOPE(-63.550,-63.550,-64.833,-64.833) |
geographic |
Galapagos Greenland Argentina Uruguay Tigre Moreno Martínez Todd Chaco Tucumán Fernandez Rodriguez Ruta Alegre Ramirez Saavedra Izquierdo Varela Entre Ríos Pergamino Daguerre Lobería Sarandí |
geographic_facet |
Galapagos Greenland Argentina Uruguay Tigre Moreno Martínez Todd Chaco Tucumán Fernandez Rodriguez Ruta Alegre Ramirez Saavedra Izquierdo Varela Entre Ríos Pergamino Daguerre Lobería Sarandí |
genre |
Greenland |
genre_facet |
Greenland |
op_relation |
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op_rights |
Open Access Creative Commons Zero v1.0 Universal https://creativecommons.org/publicdomain/zero/1.0/legalcode cc0-1.0 info:eu-repo/semantics/openAccess |
op_rightsnorm |
CC0 |
op_doi |
https://doi.org/10.5281/zenodo.5987870 https://doi.org/10.11646/zootaxa.4400.1.1 https://doi.org/10.5962/bhl.title.23819 https://doi.org/10.1002/jmor.20298 https://doi.org/10.5281/zenodo.1207689 https://doi.org/10.5281/zenodo.1207691 https://d |
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ftdatacite:10.5281/zenodo.5987870 2023-05-15T16:30:54+02:00 Ariadna boesenbergi Keyserling 1877 Giroti, André Marsola Brescovit, Antonio Domingos 2018 https://dx.doi.org/10.5281/zenodo.5987870 https://zenodo.org/record/5987870 unknown Zenodo http://zenodo.org/record/1207641 http://publication.plazi.org/id/FFA2FFA9FFE4B11A160CFFC3C759FFA5 http://zoobank.org/59CC08B8-5B13-42EC-943E-C930D7D0DD7F https://zenodo.org/communities/biosyslit https://dx.doi.org/10.11646/zootaxa.4400.1.1 http://zenodo.org/record/1207641 http://publication.plazi.org/id/FFA2FFA9FFE4B11A160CFFC3C759FFA5 https://dx.doi.org/10.5962/bhl.title.23819 https://dx.doi.org/10.1002/jmor.20298 https://dx.doi.org/10.5281/zenodo.1207689 https://dx.doi.org/10.5281/zenodo.1207691 https://dx.doi.org/10.5281/zenodo.1207787 https://dx.doi.org/10.5281/zenodo.1207643 https://dx.doi.org/10.5281/zenodo.1207679 https://dx.doi.org/10.5281/zenodo.1207685 https://dx.doi.org/10.5281/zenodo.1207709 https://dx.doi.org/10.5281/zenodo.1207737 https://dx.doi.org/10.5281/zenodo.1207741 https://dx.doi.org/10.5281/zenodo.1207747 http://zoobank.org/59CC08B8-5B13-42EC-943E-C930D7D0DD7F https://dx.doi.org/10.5281/zenodo.5987869 https://zenodo.org/communities/biosyslit Open Access Creative Commons Zero v1.0 Universal https://creativecommons.org/publicdomain/zero/1.0/legalcode cc0-1.0 info:eu-repo/semantics/openAccess CC0 Biodiversity Taxonomy Animalia Arthropoda Arachnida Araneae Segestriidae Ariadna Ariadna boesenbergi article-journal ScholarlyArticle Taxonomic treatment Text 2018 ftdatacite https://doi.org/10.5281/zenodo.5987870 https://doi.org/10.11646/zootaxa.4400.1.1 https://doi.org/10.5962/bhl.title.23819 https://doi.org/10.1002/jmor.20298 https://doi.org/10.5281/zenodo.1207689 https://doi.org/10.5281/zenodo.1207691 https://d 2022-04-01T08:54:50Z Ariadna boesenbergi Keyserling, 1877 Figures 22‒23, 70 Ariadne bösenbergii Keyserling, 1877: 223, pl. 7, fig. 7 (♀ lectotype and ♂ lectoparatype from Montevideo [34°53'01"S; 56°10'55"W], Uruguay, Bösenberg leg., deposited in ZMH 58, designated by Beatty (1970), examined; ♂ type from Uruguay, deposited in NHM 90.7.1.475-6-7, examined). Ariadna bösenbergi — Petrunkevitch (1911: 130).— Mello-Leitão (1943: 233). Ariadne bösenbergi — Gerhardt (1921: 92). Ariadna mollis— Mello-Leitão (1933: 12, Syn. with A. boesenbergi 1947: 234).— Bonnet (1955: 730, 735). Ariadna boesenbergi— Beatty (1970: 476, remove the syn. with A. mollis , descr. ♀).— Prandi (1990: 3).— Grismado (2008: 357, figs 8N, O, 10E, H).— Ramírez (2014: 211, figs 139A, 141D, 167A).— Grismado & Izquierdo (2014: 154, figs 8, 10a‒b, 12).— World Spider Catalog (2018). Additional material examined. BRAZIL: Rio Grande do Sul State: Itaqui /Maçambará, Reserva Biológica São Donato [29°02'46,7"S; 56°04'48,7"W], 1♂, 23.I.2012, M. Machado leg. (IBSP 166778); Quaraí [30°23'17"S; 56°29'56"W], Cerro do Jarau, 1♀, 23.II.1975, A.A. Lise leg. (MCN 02508); Rio Grande [32°02'06"S; 52°05'55"W], 1♀ 4imm. (NHM 89.2.17; 90.7.1.477); ARGENTINA: Tucumán Province: Arroyo India Muerta, Road to Ticucho [26°31'S; 65°15'W], 1♀, 25.VI.1995, M. Ramírez & P. Goloboff leg. (MACN-Ar 10161); La Rioja Province: Chilecito [29°10'S; 67°30'W], 1♀, 27‒30.I.1956, M.E. Galiano leg. (MACN-Ar 10163); Entre Ríos Province: Concordia [31°24'S; 58°02'W], under bark of Melia azedarach , 1♀, X.1992, C. Grismado leg. (MACN-Ar 10175); Córdoba Province: Córdoba [31°25'S; 64°11'W], Bosque Alegre, 3♀ 2imm., IX.1981, P. Goloboff leg. (MACN-Ar 10166; 10167; 10168); Nono, 880m (31°47'44.3"S; 64°00'6,26"W), 2♀ 3imm., 3‒9.II.2008, M. Ramírez leg. (MACN-Ar 13504; 13505); Leones [32°42'21"S; 62°06'30"W], 1♀, 13.X.1946, W. Patridge leg. (MACN-Ar 10164); 1♀, 12.X.1946 (MACN-Ar 10234); Santa Fé Province: Santa Fé [31°39'00"S; 60°43'60"W], 2♂, I.1965, M. Aschenbach leg. (MACN-Ar 5853); 1♀, 9.X.1963, Pignalloni leg. (MACN-Ar 10176); San Luís Province: Cortaderas (in wood of " Molles ", Schinus sp., under stones) (32°29'58,9"S; 65°00'01,5"W), 1♀, 10.XI.1982, A. Roig leg. (MACN-Ar 10169); Carolina [32°48'S; 66°06'W], between stones, 3♀, XI.1992, C. Scioscia leg. (MACN-Ar 10171; 10172); Quebrada Merlo, 3♀ 1imm., 12.XI.1982, A. Roig leg. (MACN-Ar 10170); Sierra Central, 3♀ (Ex-MNHNP-AR 9073); Buenos Aires Province: San Pedro [33°40'46"S; 59°40'01"W], in a tube under bark of Eucalyptus, 1♀, 24.XII.1912, A.G. Frers leg. (MACN-Ar 10187); Pergamino [33°44'S; 60°36'W], 2♀, IX.2007, S. Rodriguez Gil leg. (MACN-Ar 35097; 35105); Fontezuela [33°55'S; 60°28'W], 5♀, IX.2007, S. Rodriguez Gil leg. (MACN-Ar 35089; 35094; 35096; 35098; 35100); Ruta Pergamino- Arrecifes, km 210, 5♀, IX.2001, S. Rodriguez Gil leg. (MACN-Ar 28170; 35099; 35102; 35103; 35104); Arrecifes [34°02'S; 60°10'W], Todd, 1♀, IX.2007, S. Rodriguez Gil leg. (MACN-Ar 35106); Zelaya [34°21'38"S; 58°52'36"W], 2♀ 1imm., VIII.1939, H. Hepper leg. (MACN-Ar 10188; 10189; 10231); Tigre [34°25'S; 58°35'W], 1♂, I.1938, J.M. Viana leg. (MACN-Ar 303); Martínez [34°29'S; 58°30'W], 3♂ 4♀ 4imm., V.1976, R. Maynard leg. (MNHNU 1036); Saavedra [34°33'18,30"S; 58°29'32,67"W], 1♀, VIII.1937, Castillo leg. (MACN-Ar 706); Junín [34°35'38"S; 60°56'47"W], 1♀, II.1962, Daguerre leg. (MACN-Ar 10186); Buenos Aires [34°35'59"S; 58°22'55"W], Capital Federal, 7♀, 1967, Bachmann leg. (MACN-Ar 21349); 4♀, VII.1945 (MACN-Ar 1585); 2♂, I.1989, M. Miranda leg. (MACN-Ar 10177; 21308); 3♀ 1imm., VIII.1948, D.Y. Carpintero & O. Carpintero leg. (MACN-Ar 10178); 1♀, VI.1964, A. Martínez leg. (MACN-Ar 10180); 4♀, 1967, Bachmann leg. (MACN-Ar 21309); 1♂, 30.XII.2008, C. Kopuchian leg. (MACN-Ar 21312); 4♀ 1imm., Bery leg. (MNHNP-AR 7045); 1♀ 2imm., 1‒31.XII.1968, Bachmann leg. (MNHNP-AR 14916); Parque Patrícios, 1♀, X.1948 (MACN-Ar 10179); Parque Centenário, under bark of Tipuana tipu , 1♀, X.1992, C. Grismado leg. (MACN-Ar 10182); Vila Lugano, 1♀ 1imm., L.R. Rodriguez leg. (MACN-Ar 10183; 10184); bit to Paladino, 1♂, XII.1965 (MACN-Ar 10185); Arbe frente ao Museo de Ciencias Naturales, 1♀ 4imm., 9.XI.1960 (MACN-Ar 21 360); Jardines del Museo de Ciencias Naturales, 1♀, III.2010, C. Grismado & M. Izquierdo leg. (MACN-Ar 2 9449); Hurlingham, Arroyo Morón [34°36'S; 58°38'W], 2♀ 1imm., XI.1962, E. Massoia leg. (MACN-Ar 10190); Moreno [34°39'04"S; 58°47'24"W], 1♀ 1imm., III.1942, L.R.D. Schiapelli leg. (MACN-Ar 2354); 2♀ 1imm., III.1 947, Exp. R.D. Schiapelli leg. (MACN-Ar 10191); 2♀, X.1947, R.D. Schiapelli leg. (MACN-Ar 21310); Sarandí, Plaza del Viaducto [34°39'45"S; 58°21'54"W], 3♀ 34imm., 30.IX.2007, C. & A. Grismado leg. (MACN-Ar 21311; 21313; 21314); Sarandí [34°41'S; 58°20'W], on wall of bricks, 3♀ 2imm., 27.IV.2002, C., D., & A. Grismado leg. (MACN-Ar 10204); Sarandí, 1♀, 15.VII.2006, C. Grismado & A. Raya leg. (MACN-Ar 11425); Villa Madero [34°42'S; 58°30'W], 5♀, VI.2007, C. Scioscia leg. (MACN-Ar 35090; 35091; 35095; 35101; 35107); 2♀, VIII.2007 (MACN-Ar 35092; 35093); Florencio Varela [34°47'51"S; 58°16'34"W], 1♀, XII.1939, F. Monrós leg. (MACN-Ar 10197); Glew [34°53'17"S; 58°23'03"W], 1♀, 1970, Carpintero leg. (MACN-Ar 10194); 1♀ 1imm., 1979 (MACN-Ar 10196); 1imm., 1972 (MACN-Ar 10195); Tandil [37°19'43"S; 59°08'13"W], La Cascada, 1♀, 16.V.1973, C. Cesari leg. (MACN-Ar 10200); Tandil, 1♀ 3imm., V.1967, E. Maury leg. (MACN-Ar 10199); Sierra de los Padres [37°57'S; 57°46'W], 2♀ 2imm., XI.1962, M.E. Galiano leg. (MACN-Ar 10202); Tornquist, Estancia Fortin Chaco [38°06'S; 62°13'W], 1♂, 1‒31.I.1972, J.A. Obarrio leg. (MNHNP-AR 14915); 15km O de Lobería [38°08'59"S; 58°47'00"W], 1♀, 4.IX.1972 (MACN-Ar 10127); Río Negro Province: General Fernandez Oro [38°57'S; 67°55'W], 1♂, I.1976, Coscaron leg. (AMNH); URUGUAY: Without specific data: 3♀ 1imm. (MACN- Ar 21361); Cerro Largo Department: Ruta 8, Rio Tacuari, 2♀, 15.IV.1965, F. Achaval leg. (MNHNU 1050); Río Negro Department: San Javier [32°41'S; 58°08'W], 4♀, VI.1962, L.C. Zolessi leg. (MNHNU 569); Durazno Department: Durazno, Ruta 5, km 172 [33°24'47"S; 52°30'02"W], 6♀ 4imm., 15.V.2009, M. Simó y col. leg. (MNHNU FCE 3152); Lavalleja Department: Minas [34°22'39"S; 55°14'17"W], 2♀ 1imm., 5.I.1990, L. Prandi leg. (MNHNU 1070); Colonia Department: Colonia del Sacramento [34°38'05"S; 57°50'27"W], 6♂ 9♀ 1imm., 27.VIII.1989, L. Prandi leg. (MNHNU 391); San José Department: Ciudad del Plata, Playa Pascual [34°44'43"S; 56°27'07"W], 1♀ 5imm., 13.I.1990, L. Prandi leg. (MNHNU 540); Canelones Department: Parque Natural INIA Las Brujas [34°38'S; 56°20'W], 4♀, 6.VI.2013, A.D. Brescovit leg. (IBSP 166779); Montevideo Department: Montevideo, 7ª Sec. Judicial [34°53'01"S; 56°10'55"W], 1♂, VI.1964, R. Capocasale leg. (CAS 9039513); Montevideo, Buceo [34°53'59"S; 56°07'58"W], 21♀ 5imm., I.1976, R. Maynard leg. (MNHNU 575; Ex-MNHNU 656); Montevideo, Cordón [34°54'00"S; 56°10'50"W], 2♂, VI.1964, R. Capocasale leg. (MNHNU 1020). Diagnosis. Ariadna boesenbergi is similar to A. crassipalpa , A. mollis , A. calilegua , A. changellkuk, A. levii and A. perkinsi due to the dark abdominal pattern with arrow-shaped marks (Figs 1A‒C, 18A, F, 20A, F, 22A, F, 31A, E, 45A, F, 47A, F, 50A, F), but differ, except A. changellkuk , by the presence of males with metatarsi I straight, without modified macrosetae (Fig. 22C‒E); it differs from A. changellkuk by having a less elongated body (Fig. 22A‒B) and a more elongated mid-portion and embolus (Fig. 23A‒B); females are distinguished by the presence of a unillobulated anterior receptaculum, flattened laterally (Fig. 23C‒D). Description. Male (IBSP 166778). Carapace and chelicerae reddish brown (Fig. 22A). Endites brownish yellow; labium dark brownish yellow. Sternum brownish yellow, with darker margins, marbled with dark pigment, and three pairs of dark marginal areas, the hind one fainted (Fig. 22B). Palps brownish orange (Fig. 23A‒B). Legs brownish yellow, pairs I‒II darker with dark pigment forming diffuse rings in the distal region of metatarsus and tarsus (Fig. 22C‒E). Abdomen withish grey, with a conspicuous dark dorsal pattern comprising arrow-shaped marks (Fig. 22A‒B). Total length 6.64; carapace 3.4 long, 2.16 wide. Chelicerae not projected (Fig. 22A). Palps: bulb elliptical, mid-portion slightly shorter than the embolus; embolus tubular, tapered, and abruptly curved; tibia short and slightly incrassated, almost cylindrical, with the dorso-ventral diameter almost equal to the bulb (Fig. 23A‒B). Legs: robust and coated with setae (Fig. 22C‒E); tarsal brush of putative chemosensory setae present in legs I‒IV, but reduced in legs I, III and IV. Leg formula: I-II-IV-III. Leg measurements: I, femur 3.16, patella 1,.2, tibia 2.6, metatarsus 3.04, tarsus 0.8, total 10.8; II, 2.88, 1.16, 2.6, 2.8, 0.76, 10.2; III, 2.0, 0.96, 1.56, 1.84, 0.64, 7.0; IV, 2.56, 1.24, 2.04, 2.0, 0.6, 8.44. Macrosetae: I, femur D0-0-0-1-1-1/0-2-0, P0-0-0-0-0-2-1-0; tibia P0-1-0-1- 0-1-0, vp1-0-1-0-1-0-1, vr2-0-1-0-1-0-1, R1-1-1-1-1-1-0; metatarsus p0-1-0-0-0-0-0-0, vp0-1-0-0-1-0-0-1, vr0-1- 0-0-1-0-0-1, r0-1-0-0-0-0-0-0 (Fig. 22C‒E); II, femur D0-0-1-1-1-0-2-0, P0-0-0-0-0-1-1-0; tibia P0-1-0-1-0-1-0, vp1-0-1-0-1-0-1, vr2-0-1-0-1-0-1, R1-1-0/1-1-0/1-1-0; metatarsus p0-1-0-0-0-0-0-0, vp0-1-0-0-1-0-0-1, vr0-1-0-0- 1-0-0-1, r0-1-0-1-0-1-0-0; IV, femur D0/1-1-1-0-1-1-0-1-0-0-0; tibia vp0-0/1-0-0-0-0-0, vr0-0-0-0-0-0-0/1; metatarsus vp0-0-0/1-0-0-0-2, vr metatarsal comb with 4 macrosetae. Female (IBSP 166779). Carapace and chelicerae brown, with cephalic region darker (Fig. 22F). Endites brown; labium dark brown. Sternum brownish orange, with darker margins, marbled with dark pigment, and three pairs of dark marginal areas (Fig. 22G). Palps brownish yellow, with tibia and tarsus reddish. Legs brownish yellow, with pairs I‒II darker. Metatarsus and tarsus of legs I‒II darker than the other segments. Dark pigment forming diffuse rings in the distal region of tibia, metatarsus and tarsus, of legs I‒II (Fig. 22I ‒J). Abdomen as in male (Fig. 22F‒G). Total length 9.88; carapace 4.44 long, 2.68 wide. Chelicerae not projected (Fig. 22F). Legs: robust and densely coated with setae, mainly on the ventral region of pairs I‒III; legs I‒II with ventro-prolateral and ventro-retrolateral tibial macrosetae of similar size, and metatarsal macrosetae of different sizes (Fig. 22I ‒J). Leg formula: I-II-IV-III. Leg measurements: I, femur 3.04, patella 1.52, tibia 2.36, metatarsus 2.32, tarsus 0.76, total 10.0; II, 2.76, 1.48, 2.32, 2.16, 0.68, 9.4; III, 2.16, 1.24, 1.48, 1.52, 0.6, 7.0; IV, 2.8, 1.56, 2.16, 1.8, 0.64, 8.96. Macrosetae: I, femur P0-0-0-0-0-1-0; tibia vp1-0-1-0-1-0-1, vr1-0-1-0-1-0-1, R0-1-0-0-0-0-0; metatarsus vp1-1-0/ 1-1-1-1-1-1, vr1-1-1-1-1-1-1 (Fig. 22I ‒J); II, femur D0-0-0-0-0-0/2-0, P0-0-0-0-1-1-0; tibia P0-0-0-0/1-0-0/1-0, vp1-0-1-0-1-0-1, vr1-0-1-0-1-0-1; metatarsus vp1-1-0/1-1-1-1-1-1, vr0/1-1-1-1-1-1-1-1-1; IV, femur D0-1-1-0-0- 0-0; metatarsus vp0-0-0-0-0-0-1, vr metatarsal comb with 4 macrosetae. Genital external plate wide and conspicuous, occupying 1/3 of the epigastric region (Fig. 22G‒H). Internal genitalia: anterior receptaculum with the glandular plate rounded (Fig. 23C‒D). Variation. Male (n=10): Total length 5.12‒7.04; carapace 2.92‒3.52 long, 2.0‒2.48 wide; femur I 2.44‒3.4. Female (n=10): Total length 8.72‒11.64; carapace 3.84‒4.88 long, 2.44‒3.08 wide; femur I 2.64‒3.4. Distribution. South of Brazil, Uruguay, and center and east of Argentina (Fig. 70). : Published as part of Giroti, André Marsola & Brescovit, Antonio Domingos, 2018, The taxonomy of the American Ariadna Audouin (Araneae: Synspermiata: Segestriidae), pp. 1-114 in Zootaxa 4400 (1) on pages 37-41, DOI: 10.11646/zootaxa.4400.1.1, http://zenodo.org/record/1207641 : {"references": ["Keyserling, E. (1877) Amerikanische Spinnenarten aus den Familien der Pholcoidae, Scytodoidae und Dysderoidae. Verhandlungen der Zoologisch-Botanischen Gesellschaft in Wien, 27, 205 - 234.", "Beatty, J. A. (1970) The spider genus Ariadna in the Americas (Araneae, Dysderidae). Bulletin of the Museum of Comparative Zoology, 139 (8), 433 - 518.", "Petrunkevitch, A. (1911) A synonymic index-catalogue of spiders of North, Central and South America with all adjacent islands, Greenland, Bermuda, West Indies, Terra del Fuego, Galapagos, etc. Bulletin of the American Museum of Natural History, 29, 1 - 791. https: // doi. org / 10.5962 / bhl. title. 23819", "Mello-Leitao, C. F. de (1943) Catalogo das aranhas do Rio Grande do Sul. Arquivos do Museu Nacional do Rio de Janeiro, 37, 147 - 245.", "Gerhardt, U. (1921) Vergleichende Studien uber die Morphologie des mannlichen Tasters und die Biologie der Kopulation der Spinnen. Archiv fur Naturgeschichte, 87, 78 - 247.", "Mello-Leitao, C. F. de (1933) Catalogo das aranhas argentinas. Archivos da Escola Superior de Agricultura e Medicina Veterinaria, Rio de Janeiro, 10, 3 - 63.", "Bonnet, P. (1955) Bibliographia araneorum. 2 (1). Les Artisans de L'Imprimerie Douladoure, Toulouse, 919 pp.", "Prandi, L. (1990) El genero Ariadna (Araneae, Segestriidae) en la Republica Oriental del Uruguay. 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Avaliable from: http: // wsc. nmbe. ch / (accessed on 16 January 2018)"]} Text Greenland DataCite Metadata Store (German National Library of Science and Technology) Galapagos Greenland Argentina Uruguay Tigre ENVELOPE(-67.700,-67.700,-67.183,-67.183) Moreno ENVELOPE(-62.300,-62.300,-64.083,-64.083) Martínez ENVELOPE(-62.183,-62.183,-64.650,-64.650) Todd ENVELOPE(-85.933,-85.933,-78.050,-78.050) Chaco ENVELOPE(-60.583,-60.583,-63.033,-63.033) Tucumán ENVELOPE(-63.200,-63.200,-64.683,-64.683) Fernandez ENVELOPE(-62.233,-62.233,-63.250,-63.250) Rodriguez ENVELOPE(-56.720,-56.720,-63.529,-63.529) Ruta ENVELOPE(18.412,18.412,69.835,69.835) Alegre ENVELOPE(-67.050,-67.050,-68.217,-68.217) Ramirez ENVELOPE(-56.683,-56.683,-63.583,-63.583) Saavedra ENVELOPE(-57.931,-57.931,-63.317,-63.317) Izquierdo ENVELOPE(-56.950,-56.950,-64.333,-64.333) Varela ENVELOPE(-60.583,-60.583,-62.933,-62.933) Entre Ríos ENVELOPE(-66.333,-66.333,-66.433,-66.433) Pergamino ENVELOPE(-41.417,-41.417,-82.250,-82.250) Daguerre ENVELOPE(-63.417,-63.417,-65.117,-65.117) Lobería ENVELOPE(-60.800,-60.800,-62.450,-62.450) Sarandí ENVELOPE(-63.550,-63.550,-64.833,-64.833) |