Grammapsychops lebedevi Martynova 1954
Grammapsychops lebedevi Martynova, 1954 Figs 1–3 Grammapsychops lebedevi Martynova 1954: 1167 –1169, Fig. 1; Rohdendorf 1957: 40, 88, Fig. 18 [Psychopidae (sic)]; Carpenter 1992: 351, Fig. 195,1; Engel & Grimaldi 2008: 13 (Tab. 6) [Psychopsidae s. str.]; Peng et al . 2011: 120 (Tabl. 1). Grammop...
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openpolar |
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Biodiversity Taxonomy Animalia Arthropoda Insecta Neuroptera Psychopsidae Grammapsychops Grammapsychops lebedevi |
spellingShingle |
Biodiversity Taxonomy Animalia Arthropoda Insecta Neuroptera Psychopsidae Grammapsychops Grammapsychops lebedevi Makarkin, Vladimir N. Grammapsychops lebedevi Martynova 1954 |
topic_facet |
Biodiversity Taxonomy Animalia Arthropoda Insecta Neuroptera Psychopsidae Grammapsychops Grammapsychops lebedevi |
description |
Grammapsychops lebedevi Martynova, 1954 Figs 1–3 Grammapsychops lebedevi Martynova 1954: 1167 –1169, Fig. 1; Rohdendorf 1957: 40, 88, Fig. 18 [Psychopidae (sic)]; Carpenter 1992: 351, Fig. 195,1; Engel & Grimaldi 2008: 13 (Tab. 6) [Psychopsidae s. str.]; Peng et al . 2011: 120 (Tabl. 1). Grammopsychops [sic] lebedevi : Martynova 1962: Fig. 864; Kolosnitsyna 1964: 149 [Psychopsydidae (sic)]; Zherikhin 1978: 80 [Psychopsidae]. Re-description of the holotype . Hind wing ca. 41 mm long; maximum width (measured perpendicular to RA) 18.5 mm as preserved. Distal nygma absent; basal nygma not detected. Costal space strongly dilated along entire preserved length, slightly narrowed towards apex. Subcostal veinlets closely spaces, nearly all once to twice forked, becoming more oblique towards apex. Costal crossveins numerous, mainly arranged in gradate series orientated obliquely to anterior wing margin. Subcostal space narrow along entire length (its width in Fig. 3 is seen slightly narrower than in the reality as RA is convex and the space is at the angle); crossveins not preserved. ScP apically not fused with RA, entering margin well before wing apex. R broad; RA strongly convex, entering margin just before wing apex, with three long veinlets. RA (between RA and RP) space very narrow, slightly broadened distad; few distal crossveins detected. RP originates close to wing base, with 13 pectinate branches; broad proximally (up to origin of RP6). RP1 – RP6 broad proximally; RP1 – RP4 probably shallowly branched (their distal parts not preserved); RP5 – RP11 deeply dichotomously branched; RP12, RP13 dichotomously branched, but more distally. Proximal branches of RP basally with setal sockets arranged mainly in two rows (Fig. 2B); distal parts of RP branches appear thin as located in deep depressions, and setal sockets not detected. M forked slightly distad origin of RP1. MA, MP parallel, not branched for preserved proximal half (not preserved distally). Cu divided into CuA and CuP close to base of wing. CuA strongly concave, pectinately branched with nine preserved, very oblique branches (two of these deeply forked), their terminal parts not preserved. CuP with two anteriorly directed, parallel branches; their terminal parts not preserved. Anterior trace of AA1 fragmentarily preserved, close and parallel to posterior trace of CuP. Crossveins in radial to cubital spaces numerous, arranged mainly in many gradate series; distinct outer gradate series not detected; basal crossveins between R/RP and M/MA irregularly spaced. Distinct long convex fold between each vein and veinlet in their distal parts. Trichosors along costal and apical margins indistinct. Wing pattern not preserved, there are only small patches of dark pigmentation. Holotype. Specimen PIN 846 /1-2 (part only), collected by Ivan V. Lebedev in 1951. An incomplete hind wing. Type locality and horizon. Russia: Krasnoyarsk Region: District of Pirovskoe: right bank of the Kem’ River (left tributary of Yenisey River), ca. 5 km upstream from the mouth of the Belaya River [57.971372°N 92.230036°E]; exposure 0 92, horizon 3. Late Cretaceous: Cenomanian (Simonovo Formation). Remarks. Martynova (1954) interpreted this wing as the left forewing, and she believed that its posterior margin is preserved (actually, it is not preserved, possibly inwardly folded). Based on this assumption, she designated veins accordingly and incorrectly, i.e., her M is actually five proximal branches of RP; Cu (CuA and CuP) is M (MA and MP); A1 is Cu (CuA and CuP); and A2 is AA1. The venation of the holotype shows that this is a hind wing because its CuA is clearly concave. Also, M is forked very proximally and the anterior trace of AA1 is proximally parallel to the posterior trace of CuP, which are more characteristic of the hind wing than of the forewing. Martynova believed that the margins of the broad basal part of the vein R, then RP, are two parallel thin veins, i.e., R, then RP, and M in her interpretation (see Martynova 1954: Fig. 1). So, she considered ‘M’ running to the wing base as a separate vein and deeply forked into five branches (my RP1 to RP5). However, it is clearly visible that the basal parts of R and RP are broad and no break of RP between the origins of RP1 and RP6 exists (Fig. 2B). The costal space of this wing is very broad assuming that it might be a forewing. However, the costal space of the hind wing of Cretapsychops decipiens Peng et al ., 2010 is similarly configured to that of Grammapsychops lebedevi : it is very broad, wider than that of the forewing, and proximally more dilated than distally (see Peng et al . 2010: Fig. 3). Also, the costal spaces of the hind wing of Pulchroptilonia espatifata are configured rather similarly (see Martins-Neto 1997: Figs 8A, B). : Published as part of Makarkin, Vladimir N., 2018, Re-description of Grammapsychops lebedevi Martynova, 1954 (Neuroptera: Psychopsidae) with notes on the Late Cretaceous psychopsoids, pp. 581-594 in Zootaxa 4524 (5) on pages 583-585, DOI: 10.11646/zootaxa.4524.5.5, http://zenodo.org/record/2610783 : {"references": ["Martynova, O. M. (1954) Neuroptera from the Cretaceous deposits of Siberia. Doklady Akademii Nauk SSSR, 94, 1167 - 1169. [in Russian]", "Rohdendorf, B. B. (1957) Paleoentomological investigations in the USSR. Trudy Paleontologicheskogo Instituta, 66, 1 - 102. [in Russian]", "Carpenter, F. M. (1992) Superclass Hexapoda. In: Moore, R. C. & Kaesler, R. L. (Eds.), Treatise on Invertebrate Paleontology. Part R. Arthropoda 4. Fol. 4. Geological Society of America, Boulder, CO and University of Kansas, Lawrence, KS, pp. iii + 279 - 655.", "Engel, M. S. & Grimaldi, D. A. (2008) Diverse Neuropterida in Cretaceous amber, with particular reference to the paleofauna of Myanmar (Insecta). Nova Supplementa Entomologica, 20, 1 - 86.", "Peng, Y. Y., Makarkin, V. N., Wang, X. D. & Ren, D. (2011) A new fossil silky lacewing genus (Neuroptera: Psychopsidae) from the Early Cretaceous Yixian Formation of China. ZooKeys, 130, 217 - 228. https: // doi. org / 10.3897 / zookeys. 130.1576", "Martynova, O. M. (1962) Superorder Neuropteroidea. In: Rohdendorf, B. B. (Vol. Ed.), Fundamentals of Paleontology. Arthropoda-Tracheata and Chelicerata. Academy of Science of the USSR, Moscow, pp. 269 - 282. [in Russian; English translation: (1991). In: Davis, D. R. (Ed.), Fundamentals of Paleontology. Fol. 9. Arthropoda, Tracheata, Chelicerata. Smithsonian Institution Libraries and the National Science Foundation, Washington, D. C., pp. 383 - 404]", "Kolosnitsyna, G. R. (1964) New fossil insects from the Jurassic of the Irkutsk Basin]. In: Martinson, G. G. (Ed.), Stratigraphy and Paleontology of the Mesozoic and Cenozoic Deposits of Eastern Siberia and the Soviet Far East. Trudy Limnologicheskogo Instituta, 4 (24), pp. 114 - 150. [in Russian]", "Zherikhin, V. V. (1978) Development and changes of Cretaceous and Cenozoic faunistic complexes (tracheates and chelicerates). Trudy Paleontologicheskogo Instituta, 165, 1 - 200. [in Russian]", "Peng, Y. Y., Makarkin, V. N., Yang, Q. & Ren, D. (2010) A new silky lacewing (Neuroptera: Psychopsidae) from the Middle Jurassic of Inner Mongolia, China. Zootaxa, 2663, 59 - 67.", "Martins-Neto, R. G. (1997) Neuropteros (Insecta, Planipennia) da Formacao Santana (Cretaceo Inferior), Bacia do Araripe, nordeste do Brasil. X - Descricao de novos taxons (Chrysopidae, Babinskaiidae, Myrmeleontidae, Ascalaphidae e Psychopsidae). Revista Universidade de Guarulhos (Ciencias de Exatas e Technologicas), 2 (4), 68 - 83."]} |
format |
Text |
author |
Makarkin, Vladimir N. |
author_facet |
Makarkin, Vladimir N. |
author_sort |
Makarkin, Vladimir N. |
title |
Grammapsychops lebedevi Martynova 1954 |
title_short |
Grammapsychops lebedevi Martynova 1954 |
title_full |
Grammapsychops lebedevi Martynova 1954 |
title_fullStr |
Grammapsychops lebedevi Martynova 1954 |
title_full_unstemmed |
Grammapsychops lebedevi Martynova 1954 |
title_sort |
grammapsychops lebedevi martynova 1954 |
publisher |
Zenodo |
publishDate |
2018 |
url |
https://dx.doi.org/10.5281/zenodo.5954716 https://zenodo.org/record/5954716 |
long_lat |
ENVELOPE(-66.867,-66.867,-68.167,-68.167) ENVELOPE(82.680,82.680,71.828,71.828) |
geographic |
Nordeste Yenisey |
geographic_facet |
Nordeste Yenisey |
genre |
yenisey river ren Siberia |
genre_facet |
yenisey river ren Siberia |
op_relation |
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Open Access Creative Commons Zero v1.0 Universal https://creativecommons.org/publicdomain/zero/1.0/legalcode cc0-1.0 info:eu-repo/semantics/openAccess |
op_rightsnorm |
CC0 |
op_doi |
https://doi.org/10.5281/zenodo.5954716 https://doi.org/10.11646/zootaxa.4524.5.5 https://doi.org/10.5281/zenodo.2610785 https://doi.org/10.5281/zenodo.2610787 https://doi.org/10.5281/zenodo.2610789 https://doi.org/10.5281/zenodo.5954715 |
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ftdatacite:10.5281/zenodo.5954716 2023-05-15T18:45:46+02:00 Grammapsychops lebedevi Martynova 1954 Makarkin, Vladimir N. 2018 https://dx.doi.org/10.5281/zenodo.5954716 https://zenodo.org/record/5954716 unknown Zenodo http://zenodo.org/record/2610783 http://publication.plazi.org/id/FFF2F068FFB8841A731002122B65C824 http://zoobank.org/297DB9B9-F1F8-46C0-8E43-95EBA0906B9F https://zenodo.org/communities/biosyslit https://dx.doi.org/10.11646/zootaxa.4524.5.5 http://zenodo.org/record/2610783 http://publication.plazi.org/id/FFF2F068FFB8841A731002122B65C824 https://dx.doi.org/10.5281/zenodo.2610785 https://dx.doi.org/10.5281/zenodo.2610787 https://dx.doi.org/10.5281/zenodo.2610789 http://zoobank.org/297DB9B9-F1F8-46C0-8E43-95EBA0906B9F https://dx.doi.org/10.5281/zenodo.5954715 https://zenodo.org/communities/biosyslit Open Access Creative Commons Zero v1.0 Universal https://creativecommons.org/publicdomain/zero/1.0/legalcode cc0-1.0 info:eu-repo/semantics/openAccess CC0 Biodiversity Taxonomy Animalia Arthropoda Insecta Neuroptera Psychopsidae Grammapsychops Grammapsychops lebedevi article-journal ScholarlyArticle Text Taxonomic treatment 2018 ftdatacite https://doi.org/10.5281/zenodo.5954716 https://doi.org/10.11646/zootaxa.4524.5.5 https://doi.org/10.5281/zenodo.2610785 https://doi.org/10.5281/zenodo.2610787 https://doi.org/10.5281/zenodo.2610789 https://doi.org/10.5281/zenodo.5954715 2022-03-10T16:47:14Z Grammapsychops lebedevi Martynova, 1954 Figs 1–3 Grammapsychops lebedevi Martynova 1954: 1167 –1169, Fig. 1; Rohdendorf 1957: 40, 88, Fig. 18 [Psychopidae (sic)]; Carpenter 1992: 351, Fig. 195,1; Engel & Grimaldi 2008: 13 (Tab. 6) [Psychopsidae s. str.]; Peng et al . 2011: 120 (Tabl. 1). Grammopsychops [sic] lebedevi : Martynova 1962: Fig. 864; Kolosnitsyna 1964: 149 [Psychopsydidae (sic)]; Zherikhin 1978: 80 [Psychopsidae]. Re-description of the holotype . Hind wing ca. 41 mm long; maximum width (measured perpendicular to RA) 18.5 mm as preserved. Distal nygma absent; basal nygma not detected. Costal space strongly dilated along entire preserved length, slightly narrowed towards apex. Subcostal veinlets closely spaces, nearly all once to twice forked, becoming more oblique towards apex. Costal crossveins numerous, mainly arranged in gradate series orientated obliquely to anterior wing margin. Subcostal space narrow along entire length (its width in Fig. 3 is seen slightly narrower than in the reality as RA is convex and the space is at the angle); crossveins not preserved. ScP apically not fused with RA, entering margin well before wing apex. R broad; RA strongly convex, entering margin just before wing apex, with three long veinlets. RA (between RA and RP) space very narrow, slightly broadened distad; few distal crossveins detected. RP originates close to wing base, with 13 pectinate branches; broad proximally (up to origin of RP6). RP1 – RP6 broad proximally; RP1 – RP4 probably shallowly branched (their distal parts not preserved); RP5 – RP11 deeply dichotomously branched; RP12, RP13 dichotomously branched, but more distally. Proximal branches of RP basally with setal sockets arranged mainly in two rows (Fig. 2B); distal parts of RP branches appear thin as located in deep depressions, and setal sockets not detected. M forked slightly distad origin of RP1. MA, MP parallel, not branched for preserved proximal half (not preserved distally). Cu divided into CuA and CuP close to base of wing. CuA strongly concave, pectinately branched with nine preserved, very oblique branches (two of these deeply forked), their terminal parts not preserved. CuP with two anteriorly directed, parallel branches; their terminal parts not preserved. Anterior trace of AA1 fragmentarily preserved, close and parallel to posterior trace of CuP. Crossveins in radial to cubital spaces numerous, arranged mainly in many gradate series; distinct outer gradate series not detected; basal crossveins between R/RP and M/MA irregularly spaced. Distinct long convex fold between each vein and veinlet in their distal parts. Trichosors along costal and apical margins indistinct. Wing pattern not preserved, there are only small patches of dark pigmentation. Holotype. Specimen PIN 846 /1-2 (part only), collected by Ivan V. Lebedev in 1951. An incomplete hind wing. Type locality and horizon. Russia: Krasnoyarsk Region: District of Pirovskoe: right bank of the Kem’ River (left tributary of Yenisey River), ca. 5 km upstream from the mouth of the Belaya River [57.971372°N 92.230036°E]; exposure 0 92, horizon 3. Late Cretaceous: Cenomanian (Simonovo Formation). Remarks. Martynova (1954) interpreted this wing as the left forewing, and she believed that its posterior margin is preserved (actually, it is not preserved, possibly inwardly folded). Based on this assumption, she designated veins accordingly and incorrectly, i.e., her M is actually five proximal branches of RP; Cu (CuA and CuP) is M (MA and MP); A1 is Cu (CuA and CuP); and A2 is AA1. The venation of the holotype shows that this is a hind wing because its CuA is clearly concave. Also, M is forked very proximally and the anterior trace of AA1 is proximally parallel to the posterior trace of CuP, which are more characteristic of the hind wing than of the forewing. Martynova believed that the margins of the broad basal part of the vein R, then RP, are two parallel thin veins, i.e., R, then RP, and M in her interpretation (see Martynova 1954: Fig. 1). So, she considered ‘M’ running to the wing base as a separate vein and deeply forked into five branches (my RP1 to RP5). However, it is clearly visible that the basal parts of R and RP are broad and no break of RP between the origins of RP1 and RP6 exists (Fig. 2B). The costal space of this wing is very broad assuming that it might be a forewing. However, the costal space of the hind wing of Cretapsychops decipiens Peng et al ., 2010 is similarly configured to that of Grammapsychops lebedevi : it is very broad, wider than that of the forewing, and proximally more dilated than distally (see Peng et al . 2010: Fig. 3). Also, the costal spaces of the hind wing of Pulchroptilonia espatifata are configured rather similarly (see Martins-Neto 1997: Figs 8A, B). : Published as part of Makarkin, Vladimir N., 2018, Re-description of Grammapsychops lebedevi Martynova, 1954 (Neuroptera: Psychopsidae) with notes on the Late Cretaceous psychopsoids, pp. 581-594 in Zootaxa 4524 (5) on pages 583-585, DOI: 10.11646/zootaxa.4524.5.5, http://zenodo.org/record/2610783 : {"references": ["Martynova, O. M. (1954) Neuroptera from the Cretaceous deposits of Siberia. Doklady Akademii Nauk SSSR, 94, 1167 - 1169. [in Russian]", "Rohdendorf, B. B. (1957) Paleoentomological investigations in the USSR. Trudy Paleontologicheskogo Instituta, 66, 1 - 102. [in Russian]", "Carpenter, F. M. (1992) Superclass Hexapoda. In: Moore, R. C. & Kaesler, R. L. (Eds.), Treatise on Invertebrate Paleontology. Part R. Arthropoda 4. Fol. 4. Geological Society of America, Boulder, CO and University of Kansas, Lawrence, KS, pp. iii + 279 - 655.", "Engel, M. S. & Grimaldi, D. A. (2008) Diverse Neuropterida in Cretaceous amber, with particular reference to the paleofauna of Myanmar (Insecta). Nova Supplementa Entomologica, 20, 1 - 86.", "Peng, Y. Y., Makarkin, V. N., Wang, X. D. & Ren, D. (2011) A new fossil silky lacewing genus (Neuroptera: Psychopsidae) from the Early Cretaceous Yixian Formation of China. ZooKeys, 130, 217 - 228. https: // doi. org / 10.3897 / zookeys. 130.1576", "Martynova, O. M. (1962) Superorder Neuropteroidea. In: Rohdendorf, B. B. (Vol. Ed.), Fundamentals of Paleontology. Arthropoda-Tracheata and Chelicerata. Academy of Science of the USSR, Moscow, pp. 269 - 282. [in Russian; English translation: (1991). In: Davis, D. R. (Ed.), Fundamentals of Paleontology. Fol. 9. Arthropoda, Tracheata, Chelicerata. Smithsonian Institution Libraries and the National Science Foundation, Washington, D. C., pp. 383 - 404]", "Kolosnitsyna, G. R. (1964) New fossil insects from the Jurassic of the Irkutsk Basin]. In: Martinson, G. G. (Ed.), Stratigraphy and Paleontology of the Mesozoic and Cenozoic Deposits of Eastern Siberia and the Soviet Far East. Trudy Limnologicheskogo Instituta, 4 (24), pp. 114 - 150. [in Russian]", "Zherikhin, V. V. (1978) Development and changes of Cretaceous and Cenozoic faunistic complexes (tracheates and chelicerates). Trudy Paleontologicheskogo Instituta, 165, 1 - 200. [in Russian]", "Peng, Y. Y., Makarkin, V. N., Yang, Q. & Ren, D. (2010) A new silky lacewing (Neuroptera: Psychopsidae) from the Middle Jurassic of Inner Mongolia, China. Zootaxa, 2663, 59 - 67.", "Martins-Neto, R. G. (1997) Neuropteros (Insecta, Planipennia) da Formacao Santana (Cretaceo Inferior), Bacia do Araripe, nordeste do Brasil. X - Descricao de novos taxons (Chrysopidae, Babinskaiidae, Myrmeleontidae, Ascalaphidae e Psychopsidae). Revista Universidade de Guarulhos (Ciencias de Exatas e Technologicas), 2 (4), 68 - 83."]} Text yenisey river ren Siberia DataCite Metadata Store (German National Library of Science and Technology) Nordeste ENVELOPE(-66.867,-66.867,-68.167,-68.167) Yenisey ENVELOPE(82.680,82.680,71.828,71.828) |