Heyuanninae Yun, 2019, clade. nov

• Main Author: Yun, Chan-Gyu
• Format: Text
• Language: unknown
• Published: Zenodo 2019
• Subjects: Biodiversity; Taxonomy; Animalia; Chordata; Reptilia; Dinosauria; Oviraptoridae; Antarctic; Canada; Salisbury; Osborn; Currie; Tait; Dodson; Antarc*
• Online Access: https://dx.doi.org/10.5281/zenodo.5933382; https://zenodo.org/record/5933382

Barsbold (1981) erected Ingenia yanshini Barsbold, 1981 for a nearly complete oviraptorid skeleton from the Upper Cretaceous (Campanian-Maastrichtian) Barun Goyot Formation in Ömnögov, southern Mongolia. Moreover, he erected a subfamily-level clade Ingeniinae Barsbold, 1981 to distinguish Ingenia yanshini from other oviraptorids such as Oviraptor philoceratops Osborn, 1924 (Barsbold, 1981). When Barsbold first erected the subfamily, he considered this subfamily as monotypic (Barsbold, 1981) but now this subfamily comprisesmultiple taxa such as Machairasaurus leptonychus Longrich, Currie, and Dong, 2010 and Heyuannia huangi Lü, 2002. However, the genus name Ingenia is a junior homonym of the tripyloidid nematode genus Ingenia Gerlach, 1957 and according to the Article 23.1 of the International Code of Zoological Nomenclature (ICZN, 1999), Gerlach’s 1957 name has priority. Therefore, the usage of the subfamily name Ingeniinae is problematic as it is based on a junior homonym. Paul (1988) suggested a new combination Oviraptor yanshini for “ Ingenia ” yanshini but this classification has not been supported by any subsequent authors (Easter, 2013). Easter (2013) proposed a new genus name Ajancingenia Easter, 2013 for “ Ingenia ” yanshini , but subsequent phylogenetic analyses have found “ Ingenia ” yanshini and Heyuannia huangi as sister taxa (e.g., Lamanna et al. , 2014; Funston and Currie, 2016; Lü et al. , 2016; Funston et al. , 2018). Therefore, most recent studies have considered Ajancingenia as junior synonym of Heyuannia and used new combination Heyuannia yanshini for “ Ingenia ” yanshini (e.g., Funston et al. , 2018; Herne et al. , 2018). Thus, Heyuannia is the proper synonym available as a replacement name of the junior homonym. Therefore, the author proposes a new replacement subfamily name Heyuanninae clade. nov. (from Heyuannia and –inae) for “Ingeniinae”. Osmolska et al. (2004) defined “Ingeniinae” as Conchoraptor gracilis , Ingenia yanshini , their most recent common ancestor, and all descendants. Here the author revises this definition for Heyuanninae, using Heyuannia huangi as the specifier. Heyuanninae is a node-based subfamily-level clade that can be defined as the least inclusive clade containing Heyuannia huangi and Conchoraptor gracilis . : Published as part of Yun, Chan-Gyu, 2019, Heyuanninae clade nov., a replacement name for the oviraptorid subfamily " Ingeniinae " Barsbold, 1981, pp. 295-296 in Zootaxa 4671 (2) on page 295, DOI: 10.11646/zootaxa.4671.2.11, http://zenodo.org/record/3442164 : {"references": ["Barsbold, R. (1981) [Toothless carnivorous dinosaurs of Mongolia]. Joint Soviet-Mongolian Paleontological Expedition Transactions, 15, 28 - 39. [in Russian]", "Osborn, H. F. (1924) Three new Theropoda, Protoceratops zone, central Mongolia. American Museum Novitates, 144, 1 - 12.", "Longrich, N. R., Currie, P. J. & Dong, Z. - M. 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