Subspecies and Distribution. U. a. arctos Linnaeus, 1758 — Europe and W Russia. U. a. alascensis Merriam, 1896 — most of Alaska (excluding Alaska Peninsula, SE panhandle & Kodiak Island group). U. a. beringianus Middendorff, 1853 — NE Russia (Kamchatka Peninsula & N Kuril Islands northward through the Koryak Autonomous District, and along W coast of the Sea of Okhotsk). U. a. collaris Cuvier, 1824 — Russia (Siberia, from E of the Yenisey River to the Bering Sea, but excluding Kamchatka and more southern parts of the Russian Far East), N Mongolia. U. a. dalli Merriam, 1896 — SE Alaska (N of Alexander Archipelago). U. a. gyas Merriam, 1902 — Alaska peninsula. U. a. horribilis Ord, 1815 —W Canada (Yukon, North-West Territories, British Columbia & Alberta), inland W USA (extirpated from S Wyoming to Mexico). U. a. isabellinus Horsfield, 1826 — N India, Pakistan, Afghanistan, N to Kazakhstan and Mongolia (Gobi Desert). U. a. lasiotus Gray, 1867 — Russia (Southern Kuril Islands, Sakhalin, Ussuri/Amur river region of the Russian Far East), NE China, North Korea, and Japan (Hokkaido). U. a. middendorffi Merriam, 1896 — Alaska (Kodiak Island & nearby islands). U. a. pruinosus Blyth, 1853 — Tibetan Plateau, China, N Nepal. U. a. sitkensis Merriam, 1896 — SE Alaska (Alexander Archipelago & adjacent coastal area). U. a. stikeenensis Merriam, 1914 — W Canada (W British Columbia), and formerly W USA (W Washington and Oregon). U. a. syriacus Hemprich & Ehrenberg, 1828 — Middle East, from Turkey to Iran (extirpated in Syria), Caucasus mountains of Russia, Georgia, Armenia and Azerbaijan. in Ursidae

Subspecies and Distribution. U. a. arctos Linnaeus, 1758 — Europe and W Russia. U. a. alascensis Merriam, 1896 — most of Alaska (excluding Alaska Peninsula, SE panhandle & Kodiak Island group). U. a. beringianus Middendorff, 1853 — NE Russia (Kamchatka Peninsula & N Kuril Islands northward t...

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Bibliographic Details
Main Authors:	Wilson, Don E., Mittermeier, Russell A.
Format:	Still Image
Language:	unknown
Published:	Zenodo 2009
Subjects:	Biodiversity Taxonomy Animalia Chordata Mammalia Carnivora Ursidae Ursus Ursus arctos Yukon Bering Sea Okhotsk Canada British Columbia Kamchatka Peninsula Yenisey Archipelago Kamchatka Kodiak Koryak Sakhalin yenisey river Alaska Lynx Siberia
Online Access:	https://dx.doi.org/10.5281/zenodo.5714508 https://zenodo.org/record/5714508

id	ftdatacite:10.5281/zenodo.5714508
record_format	openpolar
institution	Open Polar
collection	DataCite Metadata Store (German National Library of Science and Technology)
op_collection_id	ftdatacite
language	unknown
topic	Biodiversity Taxonomy Animalia Chordata Mammalia Carnivora Ursidae Ursus Ursus arctos
spellingShingle	Biodiversity Taxonomy Animalia Chordata Mammalia Carnivora Ursidae Ursus Ursus arctos Wilson, Don E. Mittermeier, Russell A. Subspecies and Distribution. U. a. arctos Linnaeus, 1758 — Europe and W Russia. U. a. alascensis Merriam, 1896 — most of Alaska (excluding Alaska Peninsula, SE panhandle & Kodiak Island group). U. a. beringianus Middendorff, 1853 — NE Russia (Kamchatka Peninsula & N Kuril Islands northward through the Koryak Autonomous District, and along W coast of the Sea of Okhotsk). U. a. collaris Cuvier, 1824 — Russia (Siberia, from E of the Yenisey River to the Bering Sea, but excluding Kamchatka and more southern parts of the Russian Far East), N Mongolia. U. a. dalli Merriam, 1896 — SE Alaska (N of Alexander Archipelago). U. a. gyas Merriam, 1902 — Alaska peninsula. U. a. horribilis Ord, 1815 —W Canada (Yukon, North-West Territories, British Columbia & Alberta), inland W USA (extirpated from S Wyoming to Mexico). U. a. isabellinus Horsfield, 1826 — N India, Pakistan, Afghanistan, N to Kazakhstan and Mongolia (Gobi Desert). U. a. lasiotus Gray, 1867 — Russia (Southern Kuril Islands, Sakhalin, Ussuri/Amur river region of the Russian Far East), NE China, North Korea, and Japan (Hokkaido). U. a. middendorffi Merriam, 1896 — Alaska (Kodiak Island & nearby islands). U. a. pruinosus Blyth, 1853 — Tibetan Plateau, China, N Nepal. U. a. sitkensis Merriam, 1896 — SE Alaska (Alexander Archipelago & adjacent coastal area). U. a. stikeenensis Merriam, 1914 — W Canada (W British Columbia), and formerly W USA (W Washington and Oregon). U. a. syriacus Hemprich & Ehrenberg, 1828 — Middle East, from Turkey to Iran (extirpated in Syria), Caucasus mountains of Russia, Georgia, Armenia and Azerbaijan. in Ursidae
topic_facet	Biodiversity Taxonomy Animalia Chordata Mammalia Carnivora Ursidae Ursus Ursus arctos
description	Subspecies and Distribution. U. a. arctos Linnaeus, 1758 — Europe and W Russia. U. a. alascensis Merriam, 1896 — most of Alaska (excluding Alaska Peninsula, SE panhandle & Kodiak Island group). U. a. beringianus Middendorff, 1853 — NE Russia (Kamchatka Peninsula & N Kuril Islands northward through the Koryak Autonomous District, and along W coast of the Sea of Okhotsk). U. a. collaris Cuvier, 1824 — Russia (Siberia, from E of the Yenisey River to the Bering Sea, but excluding Kamchatka and more southern parts of the Russian Far East), N Mongolia. U. a. dalli Merriam, 1896 — SE Alaska (N of Alexander Archipelago). U. a. gyas Merriam, 1902 — Alaska peninsula. U. a. horribilis Ord, 1815 —W Canada (Yukon, North-West Territories, British Columbia & Alberta), inland W USA (extirpated from S Wyoming to Mexico). U. a. isabellinus Horsfield, 1826 — N India, Pakistan, Afghanistan, N to Kazakhstan and Mongolia (Gobi Desert). U. a. lasiotus Gray, 1867 — Russia (Southern Kuril Islands, Sakhalin, Ussuri/Amur river region of the Russian Far East), NE China, North Korea, and Japan (Hokkaido). U. a. middendorffi Merriam, 1896 — Alaska (Kodiak Island & nearby islands). U. a. pruinosus Blyth, 1853 — Tibetan Plateau, China, N Nepal. U. a. sitkensis Merriam, 1896 — SE Alaska (Alexander Archipelago & adjacent coastal area). U. a. stikeenensis Merriam, 1914 — W Canada (W British Columbia), and formerly W USA (W Washington and Oregon). U. a. syriacus Hemprich & Ehrenberg, 1828 — Middle East, from Turkey to Iran (extirpated in Syria), Caucasus mountains of Russia, Georgia, Armenia and Azerbaijan. : Published as part of Don E. Wilson & Russell A. Mittermeier, 2009, Ursidae, pp. 448-497 in Handbook of the Mammals of the World – Volume 1 Carnivores, Barcelona :Lynx Edicions on page 495, DOI: 10.5281/zenodo.5714493
format	Still Image
author	Wilson, Don E. Mittermeier, Russell A.
author_facet	Wilson, Don E. Mittermeier, Russell A.
author_sort	Wilson, Don E.
title	Subspecies and Distribution. U. a. arctos Linnaeus, 1758 — Europe and W Russia. U. a. alascensis Merriam, 1896 — most of Alaska (excluding Alaska Peninsula, SE panhandle & Kodiak Island group). U. a. beringianus Middendorff, 1853 — NE Russia (Kamchatka Peninsula & N Kuril Islands northward through the Koryak Autonomous District, and along W coast of the Sea of Okhotsk). U. a. collaris Cuvier, 1824 — Russia (Siberia, from E of the Yenisey River to the Bering Sea, but excluding Kamchatka and more southern parts of the Russian Far East), N Mongolia. U. a. dalli Merriam, 1896 — SE Alaska (N of Alexander Archipelago). U. a. gyas Merriam, 1902 — Alaska peninsula. U. a. horribilis Ord, 1815 —W Canada (Yukon, North-West Territories, British Columbia & Alberta), inland W USA (extirpated from S Wyoming to Mexico). U. a. isabellinus Horsfield, 1826 — N India, Pakistan, Afghanistan, N to Kazakhstan and Mongolia (Gobi Desert). U. a. lasiotus Gray, 1867 — Russia (Southern Kuril Islands, Sakhalin, Ussuri/Amur river region of the Russian Far East), NE China, North Korea, and Japan (Hokkaido). U. a. middendorffi Merriam, 1896 — Alaska (Kodiak Island & nearby islands). U. a. pruinosus Blyth, 1853 — Tibetan Plateau, China, N Nepal. U. a. sitkensis Merriam, 1896 — SE Alaska (Alexander Archipelago & adjacent coastal area). U. a. stikeenensis Merriam, 1914 — W Canada (W British Columbia), and formerly W USA (W Washington and Oregon). U. a. syriacus Hemprich & Ehrenberg, 1828 — Middle East, from Turkey to Iran (extirpated in Syria), Caucasus mountains of Russia, Georgia, Armenia and Azerbaijan. in Ursidae
title_short	Subspecies and Distribution. U. a. arctos Linnaeus, 1758 — Europe and W Russia. U. a. alascensis Merriam, 1896 — most of Alaska (excluding Alaska Peninsula, SE panhandle & Kodiak Island group). U. a. beringianus Middendorff, 1853 — NE Russia (Kamchatka Peninsula & N Kuril Islands northward through the Koryak Autonomous District, and along W coast of the Sea of Okhotsk). U. a. collaris Cuvier, 1824 — Russia (Siberia, from E of the Yenisey River to the Bering Sea, but excluding Kamchatka and more southern parts of the Russian Far East), N Mongolia. U. a. dalli Merriam, 1896 — SE Alaska (N of Alexander Archipelago). U. a. gyas Merriam, 1902 — Alaska peninsula. U. a. horribilis Ord, 1815 —W Canada (Yukon, North-West Territories, British Columbia & Alberta), inland W USA (extirpated from S Wyoming to Mexico). U. a. isabellinus Horsfield, 1826 — N India, Pakistan, Afghanistan, N to Kazakhstan and Mongolia (Gobi Desert). U. a. lasiotus Gray, 1867 — Russia (Southern Kuril Islands, Sakhalin, Ussuri/Amur river region of the Russian Far East), NE China, North Korea, and Japan (Hokkaido). U. a. middendorffi Merriam, 1896 — Alaska (Kodiak Island & nearby islands). U. a. pruinosus Blyth, 1853 — Tibetan Plateau, China, N Nepal. U. a. sitkensis Merriam, 1896 — SE Alaska (Alexander Archipelago & adjacent coastal area). U. a. stikeenensis Merriam, 1914 — W Canada (W British Columbia), and formerly W USA (W Washington and Oregon). U. a. syriacus Hemprich & Ehrenberg, 1828 — Middle East, from Turkey to Iran (extirpated in Syria), Caucasus mountains of Russia, Georgia, Armenia and Azerbaijan. in Ursidae
title_full	Subspecies and Distribution. U. a. arctos Linnaeus, 1758 — Europe and W Russia. U. a. alascensis Merriam, 1896 — most of Alaska (excluding Alaska Peninsula, SE panhandle & Kodiak Island group). U. a. beringianus Middendorff, 1853 — NE Russia (Kamchatka Peninsula & N Kuril Islands northward through the Koryak Autonomous District, and along W coast of the Sea of Okhotsk). U. a. collaris Cuvier, 1824 — Russia (Siberia, from E of the Yenisey River to the Bering Sea, but excluding Kamchatka and more southern parts of the Russian Far East), N Mongolia. U. a. dalli Merriam, 1896 — SE Alaska (N of Alexander Archipelago). U. a. gyas Merriam, 1902 — Alaska peninsula. U. a. horribilis Ord, 1815 —W Canada (Yukon, North-West Territories, British Columbia & Alberta), inland W USA (extirpated from S Wyoming to Mexico). U. a. isabellinus Horsfield, 1826 — N India, Pakistan, Afghanistan, N to Kazakhstan and Mongolia (Gobi Desert). U. a. lasiotus Gray, 1867 — Russia (Southern Kuril Islands, Sakhalin, Ussuri/Amur river region of the Russian Far East), NE China, North Korea, and Japan (Hokkaido). U. a. middendorffi Merriam, 1896 — Alaska (Kodiak Island & nearby islands). U. a. pruinosus Blyth, 1853 — Tibetan Plateau, China, N Nepal. U. a. sitkensis Merriam, 1896 — SE Alaska (Alexander Archipelago & adjacent coastal area). U. a. stikeenensis Merriam, 1914 — W Canada (W British Columbia), and formerly W USA (W Washington and Oregon). U. a. syriacus Hemprich & Ehrenberg, 1828 — Middle East, from Turkey to Iran (extirpated in Syria), Caucasus mountains of Russia, Georgia, Armenia and Azerbaijan. in Ursidae
title_fullStr	Subspecies and Distribution. U. a. arctos Linnaeus, 1758 — Europe and W Russia. U. a. alascensis Merriam, 1896 — most of Alaska (excluding Alaska Peninsula, SE panhandle & Kodiak Island group). U. a. beringianus Middendorff, 1853 — NE Russia (Kamchatka Peninsula & N Kuril Islands northward through the Koryak Autonomous District, and along W coast of the Sea of Okhotsk). U. a. collaris Cuvier, 1824 — Russia (Siberia, from E of the Yenisey River to the Bering Sea, but excluding Kamchatka and more southern parts of the Russian Far East), N Mongolia. U. a. dalli Merriam, 1896 — SE Alaska (N of Alexander Archipelago). U. a. gyas Merriam, 1902 — Alaska peninsula. U. a. horribilis Ord, 1815 —W Canada (Yukon, North-West Territories, British Columbia & Alberta), inland W USA (extirpated from S Wyoming to Mexico). U. a. isabellinus Horsfield, 1826 — N India, Pakistan, Afghanistan, N to Kazakhstan and Mongolia (Gobi Desert). U. a. lasiotus Gray, 1867 — Russia (Southern Kuril Islands, Sakhalin, Ussuri/Amur river region of the Russian Far East), NE China, North Korea, and Japan (Hokkaido). U. a. middendorffi Merriam, 1896 — Alaska (Kodiak Island & nearby islands). U. a. pruinosus Blyth, 1853 — Tibetan Plateau, China, N Nepal. U. a. sitkensis Merriam, 1896 — SE Alaska (Alexander Archipelago & adjacent coastal area). U. a. stikeenensis Merriam, 1914 — W Canada (W British Columbia), and formerly W USA (W Washington and Oregon). U. a. syriacus Hemprich & Ehrenberg, 1828 — Middle East, from Turkey to Iran (extirpated in Syria), Caucasus mountains of Russia, Georgia, Armenia and Azerbaijan. in Ursidae
title_full_unstemmed	Subspecies and Distribution. U. a. arctos Linnaeus, 1758 — Europe and W Russia. U. a. alascensis Merriam, 1896 — most of Alaska (excluding Alaska Peninsula, SE panhandle & Kodiak Island group). U. a. beringianus Middendorff, 1853 — NE Russia (Kamchatka Peninsula & N Kuril Islands northward through the Koryak Autonomous District, and along W coast of the Sea of Okhotsk). U. a. collaris Cuvier, 1824 — Russia (Siberia, from E of the Yenisey River to the Bering Sea, but excluding Kamchatka and more southern parts of the Russian Far East), N Mongolia. U. a. dalli Merriam, 1896 — SE Alaska (N of Alexander Archipelago). U. a. gyas Merriam, 1902 — Alaska peninsula. U. a. horribilis Ord, 1815 —W Canada (Yukon, North-West Territories, British Columbia & Alberta), inland W USA (extirpated from S Wyoming to Mexico). U. a. isabellinus Horsfield, 1826 — N India, Pakistan, Afghanistan, N to Kazakhstan and Mongolia (Gobi Desert). U. a. lasiotus Gray, 1867 — Russia (Southern Kuril Islands, Sakhalin, Ussuri/Amur river region of the Russian Far East), NE China, North Korea, and Japan (Hokkaido). U. a. middendorffi Merriam, 1896 — Alaska (Kodiak Island & nearby islands). U. a. pruinosus Blyth, 1853 — Tibetan Plateau, China, N Nepal. U. a. sitkensis Merriam, 1896 — SE Alaska (Alexander Archipelago & adjacent coastal area). U. a. stikeenensis Merriam, 1914 — W Canada (W British Columbia), and formerly W USA (W Washington and Oregon). U. a. syriacus Hemprich & Ehrenberg, 1828 — Middle East, from Turkey to Iran (extirpated in Syria), Caucasus mountains of Russia, Georgia, Armenia and Azerbaijan. in Ursidae
title_sort	subspecies and distribution. u. a. arctos linnaeus, 1758 — europe and w russia. u. a. alascensis merriam, 1896 — most of alaska (excluding alaska peninsula, se panhandle & kodiak island group). u. a. beringianus middendorff, 1853 — ne russia (kamchatka peninsula & n kuril islands northward through the koryak autonomous district, and along w coast of the sea of okhotsk). u. a. collaris cuvier, 1824 — russia (siberia, from e of the yenisey river to the bering sea, but excluding kamchatka and more southern parts of the russian far east), n mongolia. u. a. dalli merriam, 1896 — se alaska (n of alexander archipelago). u. a. gyas merriam, 1902 — alaska peninsula. u. a. horribilis ord, 1815 —w canada (yukon, north-west territories, british columbia & alberta), inland w usa (extirpated from s wyoming to mexico). u. a. isabellinus horsfield, 1826 — n india, pakistan, afghanistan, n to kazakhstan and mongolia (gobi desert). u. a. lasiotus gray, 1867 — russia (southern kuril islands, sakhalin, ussuri/amur river region of the russian far east), ne china, north korea, and japan (hokkaido). u. a. middendorffi merriam, 1896 — alaska (kodiak island & nearby islands). u. a. pruinosus blyth, 1853 — tibetan plateau, china, n nepal. u. a. sitkensis merriam, 1896 — se alaska (alexander archipelago & adjacent coastal area). u. a. stikeenensis merriam, 1914 — w canada (w british columbia), and formerly w usa (w washington and oregon). u. a. syriacus hemprich & ehrenberg, 1828 — middle east, from turkey to iran (extirpated in syria), caucasus mountains of russia, georgia, armenia and azerbaijan. in ursidae
publisher	Zenodo
publishDate	2009
url	https://dx.doi.org/10.5281/zenodo.5714508 https://zenodo.org/record/5714508
long_lat	ENVELOPE(-125.003,-125.003,54.000,54.000) ENVELOPE(160.000,160.000,56.000,56.000) ENVELOPE(82.680,82.680,71.828,71.828)
geographic	Yukon Bering Sea Okhotsk Canada British Columbia Kamchatka Peninsula Yenisey
geographic_facet	Yukon Bering Sea Okhotsk Canada British Columbia Kamchatka Peninsula Yenisey
genre	Archipelago Bering Sea Kamchatka Kamchatka Peninsula Kodiak Koryak Sakhalin Ursus arctos yenisey river Alaska Lynx Siberia Yukon
genre_facet	Archipelago Bering Sea Kamchatka Kamchatka Peninsula Kodiak Koryak Sakhalin Ursus arctos yenisey river Alaska Lynx Siberia Yukon
op_relation	http://publication.plazi.org/id/FFA4FFECF66BC766950C7F53FF82FF9D http://treatment.plazi.org/id/039D8794F661C76D95AF7E5EFDF7F970 https://zenodo.org/communities/biosyslit https://dx.doi.org/10.5281/zenodo.5714493 http://publication.plazi.org/id/FFA4FFECF66BC766950C7F53FF82FF9D https://dx.doi.org/10.5281/zenodo.5714781 http://treatment.plazi.org/id/039D8794F661C76D95AF7E5EFDF7F970 https://dx.doi.org/10.5281/zenodo.5714509 https://zenodo.org/communities/biosyslit
op_rights	Open Access info:eu-repo/semantics/openAccess
op_doi	https://doi.org/10.5281/zenodo.5714508 https://doi.org/10.5281/zenodo.5714493 https://doi.org/10.5281/zenodo.5714781 https://doi.org/10.5281/zenodo.5714509
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spelling	ftdatacite:10.5281/zenodo.5714508 2023-05-15T14:17:55+02:00 Subspecies and Distribution. U. a. arctos Linnaeus, 1758 — Europe and W Russia. U. a. alascensis Merriam, 1896 — most of Alaska (excluding Alaska Peninsula, SE panhandle & Kodiak Island group). U. a. beringianus Middendorff, 1853 — NE Russia (Kamchatka Peninsula & N Kuril Islands northward through the Koryak Autonomous District, and along W coast of the Sea of Okhotsk). U. a. collaris Cuvier, 1824 — Russia (Siberia, from E of the Yenisey River to the Bering Sea, but excluding Kamchatka and more southern parts of the Russian Far East), N Mongolia. U. a. dalli Merriam, 1896 — SE Alaska (N of Alexander Archipelago). U. a. gyas Merriam, 1902 — Alaska peninsula. U. a. horribilis Ord, 1815 —W Canada (Yukon, North-West Territories, British Columbia & Alberta), inland W USA (extirpated from S Wyoming to Mexico). U. a. isabellinus Horsfield, 1826 — N India, Pakistan, Afghanistan, N to Kazakhstan and Mongolia (Gobi Desert). U. a. lasiotus Gray, 1867 — Russia (Southern Kuril Islands, Sakhalin, Ussuri/Amur river region of the Russian Far East), NE China, North Korea, and Japan (Hokkaido). U. a. middendorffi Merriam, 1896 — Alaska (Kodiak Island & nearby islands). U. a. pruinosus Blyth, 1853 — Tibetan Plateau, China, N Nepal. U. a. sitkensis Merriam, 1896 — SE Alaska (Alexander Archipelago & adjacent coastal area). U. a. stikeenensis Merriam, 1914 — W Canada (W British Columbia), and formerly W USA (W Washington and Oregon). U. a. syriacus Hemprich & Ehrenberg, 1828 — Middle East, from Turkey to Iran (extirpated in Syria), Caucasus mountains of Russia, Georgia, Armenia and Azerbaijan. in Ursidae Wilson, Don E. Mittermeier, Russell A. 2009 https://dx.doi.org/10.5281/zenodo.5714508 https://zenodo.org/record/5714508 unknown Zenodo http://publication.plazi.org/id/FFA4FFECF66BC766950C7F53FF82FF9D http://treatment.plazi.org/id/039D8794F661C76D95AF7E5EFDF7F970 https://zenodo.org/communities/biosyslit https://dx.doi.org/10.5281/zenodo.5714493 http://publication.plazi.org/id/FFA4FFECF66BC766950C7F53FF82FF9D https://dx.doi.org/10.5281/zenodo.5714781 http://treatment.plazi.org/id/039D8794F661C76D95AF7E5EFDF7F970 https://dx.doi.org/10.5281/zenodo.5714509 https://zenodo.org/communities/biosyslit Open Access info:eu-repo/semantics/openAccess Biodiversity Taxonomy Animalia Chordata Mammalia Carnivora Ursidae Ursus Ursus arctos Image ImageObject Figure graphic 2009 ftdatacite https://doi.org/10.5281/zenodo.5714508 https://doi.org/10.5281/zenodo.5714493 https://doi.org/10.5281/zenodo.5714781 https://doi.org/10.5281/zenodo.5714509 2022-04-01T17:33:55Z Subspecies and Distribution. U. a. arctos Linnaeus, 1758 — Europe and W Russia. U. a. alascensis Merriam, 1896 — most of Alaska (excluding Alaska Peninsula, SE panhandle & Kodiak Island group). U. a. beringianus Middendorff, 1853 — NE Russia (Kamchatka Peninsula & N Kuril Islands northward through the Koryak Autonomous District, and along W coast of the Sea of Okhotsk). U. a. collaris Cuvier, 1824 — Russia (Siberia, from E of the Yenisey River to the Bering Sea, but excluding Kamchatka and more southern parts of the Russian Far East), N Mongolia. U. a. dalli Merriam, 1896 — SE Alaska (N of Alexander Archipelago). U. a. gyas Merriam, 1902 — Alaska peninsula. U. a. horribilis Ord, 1815 —W Canada (Yukon, North-West Territories, British Columbia & Alberta), inland W USA (extirpated from S Wyoming to Mexico). U. a. isabellinus Horsfield, 1826 — N India, Pakistan, Afghanistan, N to Kazakhstan and Mongolia (Gobi Desert). U. a. lasiotus Gray, 1867 — Russia (Southern Kuril Islands, Sakhalin, Ussuri/Amur river region of the Russian Far East), NE China, North Korea, and Japan (Hokkaido). U. a. middendorffi Merriam, 1896 — Alaska (Kodiak Island & nearby islands). U. a. pruinosus Blyth, 1853 — Tibetan Plateau, China, N Nepal. U. a. sitkensis Merriam, 1896 — SE Alaska (Alexander Archipelago & adjacent coastal area). U. a. stikeenensis Merriam, 1914 — W Canada (W British Columbia), and formerly W USA (W Washington and Oregon). U. a. syriacus Hemprich & Ehrenberg, 1828 — Middle East, from Turkey to Iran (extirpated in Syria), Caucasus mountains of Russia, Georgia, Armenia and Azerbaijan. : Published as part of Don E. Wilson & Russell A. Mittermeier, 2009, Ursidae, pp. 448-497 in Handbook of the Mammals of the World – Volume 1 Carnivores, Barcelona :Lynx Edicions on page 495, DOI: 10.5281/zenodo.5714493 Still Image Archipelago Bering Sea Kamchatka Kamchatka Peninsula Kodiak Koryak Sakhalin Ursus arctos yenisey river Alaska Lynx Siberia Yukon DataCite Metadata Store (German National Library of Science and Technology) Yukon Bering Sea Okhotsk Canada British Columbia ENVELOPE(-125.003,-125.003,54.000,54.000) Kamchatka Peninsula ENVELOPE(160.000,160.000,56.000,56.000) Yenisey ENVELOPE(82.680,82.680,71.828,71.828)

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