Praearchitipula apprima Gao, Shih, Kopeć, Krzemiński & Ren, 2015, sp. nov.

• Main Authors: Gao, Jiaqi, Shih, Chungkun, Kopeć, Katarzyna, Krzemiński, Wiesław, Ren, Dong
• Format: Text
• Language: unknown
• Published: Zenodo 2015
• Subjects: Biodiversity; Taxonomy; Animalia; Arthropoda; Insecta; Diptera; Limoniidae; Praearchitipula; Praearchitipula apprima; ren; Siberia
• Online Access: https://dx.doi.org/10.5281/zenodo.5690559; https://zenodo.org/record/5690559

Praearchitipula apprima sp. nov. (Figs. 3, 4) Diagnosis. Cross-vein sc-r far proximal of the origin of Rs, the distance less than one half of Rs; r-m near midlength of R 4 + 5; M 1 + 2 only 1 / 4 as long as M 1; d-cell about 1 / 9 of the length of wing. (Note: The position of r-m is unique for Praearchitipula and distinguishes this species from all other species of this genus.) Etymology. The specific name is derived from the Latin “ apprima ”, referring to the first time that a fossil pediciid is described from China. Description. A large-sized crane fly with well-preserved wings, body and head (Figs. 3 A, 3 B, 4 A). Body length 11.4–13 mm, head 0.5–0.7 mm, subcircular. The compound eyes with distinct facets, but no setae on eyes preserved. Antenna, 1.6 mm long, 2.1 times as long as the head length; left antenna with 14 flagellomeres preserved, right antenna 16 -segmented. Palpi with four visible segments, 1 mm long, 1.3 times as long as the head length. Thorax length 1.7–2.1 mm; prothorax barely visible; anterior part of scutum in mesothorax arched convex; the boundary between prescutum, scutum and scutellum distinct; mediotergite well-developed in lateral view. The legs long and slender. The abdomen well-preserved, relatively long and slender, length 8.9–10.3 mm, with 9 clear segments. Female with ovipositor preserved. Wing: The left wing of the holotype well-preserved, but part of outer posterior margin missing (Fig. 3 A, C). Length 10.8 mm and width 3.5 mm, 3.1 times as long as wide. Wing without any dark marks except for elongated pterostigma. Sc long, 0.74 times of the wing length and terminating significantly distad of the forking of R 4 + 5; cross-vein sc-r at 0.45 times of length of Sc. Rs of moderate length, strongly arcuated at origin; Rs arising from 0.45 times of length of wing, bifurcating into R 2 + 3 and R 4 + 5 at 0.66 times of length of wing; R 1 long; r-r (R 2) close to R 1 tip, transverse; R 4 + 5 short, 1 / 5 times as long as R 2 + 3; stem of vein M bifurcating at the same level of branching of Rs; cell m 1 petiolate; mM 1 + 2 1.4 times as long as dM 1 + 2; dM 1 + 2 half as long as cell m 1; r-m at 0.4 times of R 4 + 5; d-cell broad distally, 1 / 9 times length of wing; bM 3 + 4 2.2 times as long as dM 3 + 4. Cu distinctly bending at m-cu, straight beyond it. Anal veins elongate and divergent. Remarks. The current generic-level classification of recent Pediciidae is confusing and does not offer an easy solution to assign P. apprima sp. nov. to another grouping. The new species can be referred to the genus Praearchitipula based on the following characters: Rs bifurcating into R 2 + 3 and R 4 + 5; R 4 + 5 short; r-m between R 4 + 5 and M 1 + 2. In this situation, we decide to place this new species in the genus Praearchitipula temporarily. This new species most closely resembles P. trinervis Kalugina, 1985, but can be easily distinguished from the latter in having smaller body; cross-vein r-m between R 4 + 5 and M 1 + 2; cell m 1 4 / 3 times as long as dM 1 + 2. Material examined. Holotype, female, CNU-DIP-NN 2014001, a well-preserved body with wings. Paratype, female, CNU-DIP-NN 2014002. : Published as part of Gao, Jiaqi, Shih, Chungkun, Kopeć, Katarzyna, Krzemiński, Wiesław & Ren, Dong, 2015, New species and revisions of Pediciidae (Diptera) from the Middle Jurassic of northeastern China and Russia, pp. 77-86 in Zootaxa 3964 (1) on pages 80-84, DOI: 10.11646/zootaxa.3964.1.4, http://zenodo.org/record/288700 : {"references": ["Kalugina, N. S. & Kovalev, V. G. (1985) Dipterous insects of Jurassic Siberia. Paleontological Institute, Akademia Nauk, Moscow, 198 pp. [in Russian]"]}