Drepanoistodus basiovalis

Drepanoistodus basiovalis (Sergeeva, 1963) Fig. 6A–D Oistodus basiovalis Sergeeva, 1963: 96, pl. 7 figs 6–7, text-fig. 3. Drepanoistodus basiovalis – Lindström 1971: 43, text-figs 6, 8. — Stouge & Bagnoli 1990: 15, pl. 5 figs 18–24. — Dzik 1990: fig. 12; 1994: 78, pl. 16 figs. 16–20, text-fig. 1...

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Main Authors: Rasmussen, Jan Audun, Eriksson, Mats E., Lindskog, Anders
Format: Text
Language:unknown
Published: Zenodo 2021
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Online Access:https://dx.doi.org/10.5281/zenodo.5558708
https://zenodo.org/record/5558708
id ftdatacite:10.5281/zenodo.5558708
record_format openpolar
institution Open Polar
collection DataCite Metadata Store (German National Library of Science and Technology)
op_collection_id ftdatacite
language unknown
topic Biodiversity
Taxonomy
Animalia
Chordata
Conodonta
Distacodontina
Drepanoistodontidae
Drepanoistodus
Drepanoistodus basiovalis
spellingShingle Biodiversity
Taxonomy
Animalia
Chordata
Conodonta
Distacodontina
Drepanoistodontidae
Drepanoistodus
Drepanoistodus basiovalis
Rasmussen, Jan Audun
Eriksson, Mats E.
Lindskog, Anders
Drepanoistodus basiovalis
topic_facet Biodiversity
Taxonomy
Animalia
Chordata
Conodonta
Distacodontina
Drepanoistodontidae
Drepanoistodus
Drepanoistodus basiovalis
description Drepanoistodus basiovalis (Sergeeva, 1963) Fig. 6A–D Oistodus basiovalis Sergeeva, 1963: 96, pl. 7 figs 6–7, text-fig. 3. Drepanoistodus basiovalis – Lindström 1971: 43, text-figs 6, 8. — Stouge & Bagnoli 1990: 15, pl. 5 figs 18–24. — Dzik 1990: fig. 12; 1994: 78, pl. 16 figs. 16–20, text-fig. 12a; 2020: fig. 7A–E. — Rasmussen 1991: 277, fig. 6l; 2001: 71–73, pl. 5: 9 (cum. syn.). — Löfgren 1994: fig. 6.30; 2000a: fig. 4w; 2006: figs 3n, 3ab. — Viira et al . 2001: fig. 5z. — Zhen & Percival 2004: 93, fig. 11a–j. — Tolmacheva et al . 2013: pl. 3, fig. 24. partim Drepanoistodus basiovalis – Löfgren 1978: 55–56, pl. 1 figs 11–16 (only), non 17 (= D. contractus (Lindström, 1955)). — Olgun 1987: 49, pl. 6w (only). — Landing et al . 2003: fig. 4e (only). — Zhen et al . 2011: 222–227, fig. 12a?, b–n, p–q (only). cf. Drepanoistodus basiovalis – Zhang 1998: 61–62, pl. 5 figs 5–12 (unusually short upper margin of the base). ? Drepanoistodus basiovalis – Lehnert et al. 1998: 55, pl. 3 figs 6, 12 (12 may belong to Paroistodus originalis (Sergeeva, 1963)). — Boncheva et al . 2009: text-fig. 3.8 (broken element). — Albanesi & Ortega 2016: fig. 7(6) (shares characters with D. basiovalis and D . cf. balticus ). — Feltes et al . 2016: fig. 3ac. — Wu et al . 2018: fig. 5e (unusually long base compared to the cusp). non Drepanoistodus basiovalis – Gutiérrez-Marco et al . 2008: 153, figs 3.29–3.31 (may be Drepanoistodus cf. basiovalis or Drepanoistodus cf. suberectus (Branson & Mehl, 1933). — Hints et al . 2012: fig. 6h (= Drepanoistodus cf. suberectus ). — Wu et al . 2017: fig. 7u (= Drepanoistodus contractus (Lindström)). — Lindskog et al . 2020: fig. 7v–w (= Drepanoistodus viirae sp. nov.). Original diagnosis (translated from Sergeeva, 1963 [in Russian]) Inclined conodonts, almost symmetrical, with a wide shortened base, the edge of which is rounded. Material examined 33 geniculate elements including 24 from the Lynna section. Original description, slightly shortened (translated from Sergeeva, 1963 [in Russian]) Medium-sized conodonts (0.52–0.92 mm), inclined; the degree of inclination of the cusp is 45–60°, sometimes up to 80°. Base high, not very long, elongated along the CD; base length 2.5–3 times its height ( comment by the authors: “ we find the meaning of the latter measure ambiguous ”). Base wall slightly transparent near the edge, rounded. The angle between the sides AC is more than 90°; angle between AD 40–45°; corners are smoothly obtuse. Transverse in cross section, the base is oval, elongated along CD and compressed along L1L2. From the C side, the base is compressed, sometimes with a thin keel near the tip, with a small keel on side D. The sides of the base L1 and L2 are smooth and flat. Basal cavity is not always visible, it is wide, but not deep, without visible tops. The cusp is long, straight or slightly curved towards L1, sharply tapering towards the tip; compressed. The sides of the cusp are almost flat, with a welldeveloped longitudinal, wide carina on L1 and less developed carina on the side L2. The carinae usually run from the base to the tip of the cusp. Thin keels occur on the lower (D) and upper (C) parts of the cusp. Remarks In her original diagnosis, Sergeeva (1963) only included geniculate elements with a rounded basal margin in “ Oistodus ” basiovalis , which is also evident from the species epithet: basiovalis (meaning oval base). This interpretation of the geniculate element in Drepanoistodus basiovalis is followed here. Additional typical characters that may be added to the original species description include: anterior margin and upper anterior corner rounded or weakly rounded; cusp usually straight; a median or median to lower, longitudinal carina present on the inner (sometimes slightly concave) side of the element. Carina is more distinct in Darriwilian specimens than in Dapingian ones. Whereas angle A (Fig. 3) between the cusp and the upper margin is 29.6° with a standard deviation at 5.6, the mean ratio between the length of the free upper margin and free cusp (b/c ratio) reaches 0.40 with a standard deviation of 0.1 (Fig. 5B). Occurrence Drepanoistodus basiovalis occurs from the L. antivariabilis Zone (sample LY12-9) to the interzone (“uncertain interval”) between the L. variabilis Zone and the Y. crassus Zone sensu Lindskog et al. (2020) in the Lynna River section (sample LY12-21b; between LY12-21 and LY12-22). In addition, D. basiovalis has been reported from several other localities in Baltoscandia and Poland, and also outside the Baltica palaeocontinent, e.g., New Brunswick, Argentina, Australia and China (for references, see the synonymy list above). : Published as part of Rasmussen, Jan Audun, Eriksson, Mats E. & Lindskog, Anders, 2021, Middle Ordovician Drepanoistodus (Vertebrata, Conodonta) from Baltica, with description of three new species, pp. 106-134 in European Journal of Taxonomy 774 on pages 118-119, DOI: 10.5852/ejt.2021.774.1533, http://zenodo.org/record/5557148 : {"references": ["Sergeeva S. P. 1963. [Conodonts from the Lower Ordovician in the Leningrad region.] Paleontologicheshij Zhurnal 1963 (2): 93 - 108. [In Russian.]", "Lindstrom M. 1971. Lower Ordovician conodonts of Europe. Geological Society of America Memoirs 127: 21 - 61. https: // doi. org / 10.1130 / MEM 127 - p 21", "Stouge S. & Bagnoli G. 1990. Lower Ordovician (Volkhovian - Kundan) conodonts from Hagudden, northern Oland, Sweden. Palaeontographica Italica 77: 1 - 54.", "Dzik J. 1990. Conodont evolution in high latitudes of the Ordovician. Courier Forschungsinstitut Senckenberg 117: l - 28.", "Rasmussen J. A. 1991. Conodont stratigraphy of the Lower Ordovician Huk Formation at Slemmestad, southern Norway. Norsk Geologisk Tidsskrift 71: 265 - 88.", "Lofgren A. 1994. Arenig (Lower Ordovician) conodonts and biozonation in the eastern Siljan district, central Sweden. Journal of Paleontology 68: 1350 - 1368. https: // doi. org / 10.1017 / S 0022336000034338", "Viira V., Lofgren A., Magi S. & Wickstrom J. 2001. An Early to Middle Ordovician succession of conodont faunas at Maekalda, northern Estonia. Geological Magazine 138: 699 - 718. https: // doi. org / 10.1017 / S 0016756801005945", "Zhen Y. Y. & Percival I. G. 2004. Middle Ordovician (Darriwilian) conodonts from allochthonous limestones in the Oakdale Formation of central New South Wales. Alcheringa 28: 77 - 111. https: // doi. org / 10.1080 / 03115510408619276", "Tolmacheva T. Yu., Zaitsev A. V. & Alekseev A. S. 2013. Middle and Upper Ordovician conodonts of the Moscow Syneclise: new data on stratigraphy of the borehole Gavrilov Yam- 1 section. Stratigrafiya. Geologicheskaya Korrelyatsiya 21: 52 - 77. https: // doi. org / 10.1134 / S 0869593813040096", "Lofgren A. 1978. Arenigian and Llanvirnian conodonts from Jamtland, northern Sweden. Fossils and Strata 13: 1 - 129.", "Olgun O. 1987. Komponenten-Analyse und Conodonten-Stratigraphie der Orthoceratenkalksteine im Gebiet Falbygden, Vastergotland, Mittelschweden. Sveriges Geologiska Undersokning, Ser. Ca 70: 1 - 78.", "Landing E., Westrop S. R. & Kim D. H. 2003. First Middle Ordovician biota from southern New Brunswick: stratigraphic and tectonic implications for the evolution of the Avalon continent. Canadian Journal of Earth Sciences 40: 715 - 730. https: // doi. org / 10.1139 / E 03 - 009", "Zhen Y. Y., Wang Z. H., Zhang Y. D., Bergstrom S. M., Percival I. G. & Chen J. F. 2011. Middle to Late Ordovician (Darriwilian - Sandbian) conodonts from the Dawangou Section, Kalpin area of the Tarim Basin, northwestern China. Records of the Australian Museum 63: 203 - 266. https: // doi. org / 10.3853 / j. 0067 - 1975.63.2011.1586", "Zhang J. 1998. Conodonts from the Guniutan Formation (Llanvirnian) in Hubei and Hunan Provinces, south-central China. Stockholm Contributions in Geology 46: 1 - 161.", "Lehnert D., Keller M. & Bordonaro D. 1998. Early Ordovician conodonts from the southern Cuyania terrane (Mendoza Province, Argentina). In: H. Szaniawski (ed.) Proceedings of the Sixth European Conodont Symposium (ECOS VI). Palaeontologia Polonica 58: 47 - 65.", "Boncheva I., Goncuoglu M. C., Leslie S. A., Lakova I., Sachanski V., Saydam G., Gedik I. & Koenigshof P. 2009. New conodont and palynological data from the Lower Palaeozoic in Northern Camdag, NW Anatolia, Turkey. Acta Geologica Polonica 59: 157 - 171.", "Albanesi G. L. & Ortega G. 2016. Conodont and graptolite biostratigraphy of the Ordovician System of Argentina. In: Montenari M. (ed.) Stratigraphy and Timescales 1: 61 - 121. https: // doi. org / 10.1016 / bs. sats. 2016.10.002", "Feltes N. A., Albanesi G. L. & Bergstrom S. M. 2016. Conodont biostratigraphy and global correlation of the Middle Darriwilian - Lower Sandbian Las Aguaditas Formation, Precordillera of San Juan, Argentina. Andean Geology 43: 60 - 85. https: // doi. org / 10.5027 / andgeoV 43 n 1 - a 04", "Wu R. - C., Calner M., Lehnert O., Lindskog A. & Joachimski M. 2018. Conodont biostratigraphy and carbon isotope stratigraphy of the Middle Ordovician (Darriwilian) Komstad Limestone, southern Sweden. GFF 140: 44 - 54. https: // doi. org / 10.1080 / 11035897.2018.1435561", "Gutierrez-Marco J. C., Albanesi G. L., Sarmiento G. N. & Carlotto V. 2008. An Early Ordovician (Floian) Conodont Fauna from the Eastern Cordillera of Peru (Central Andean Basin). Geologica Acta 6: 147 - 160.", "Branson E. B. & Mehl M. G. 1933. Conodont studies no. 2; conodonts from Joachim (Middle Ordovician) of Missouri; from the Plattin (Middle Ordovician) of Missouri; from the Maquoketa-Thebes (Upper Ordovician) of Missouri; a study of Hinde's types of conodonts preserved in the British Museum. Missouri University Studies 8: 77 - 167.", "Hints O., Viira V. & Nolvak J. 2012. Darriwilian (Middle Ordovician) conodont biostratigraphy in NW Estonia. Estonian Journal of Earth Sciences 61: 210 - 226. https: // doi. org / 10.3176 / earth. 2012.4.03", "Wu R. - C., Calner M. & Lehnert O. 2017. Integrated conodont biostratigraphy and carbon isotope chemostratigraphy in the Lower-Middle Ordovician of southern Sweden reveals a complete record of the MDICE. Geological Magazine 154: 334 - 353. https: // doi. org / 10.1017 / S 0016756816000017", "Lindskog A., Eriksson M. E., Rasmussen J. A., Dronov A. & Rasmussen C. M. O. 2020. Middle Ordovician carbonate facies development, conodont biostratigraphy and faunal diversity patterns at the Lynna River, northwestern Russia. Estonian Journal of Earth Sciences 69: 37 - 61. https: // doi. org / 10.3176 / earth. 2020.1.03"]}
format Text
author Rasmussen, Jan Audun
Eriksson, Mats E.
Lindskog, Anders
author_facet Rasmussen, Jan Audun
Eriksson, Mats E.
Lindskog, Anders
author_sort Rasmussen, Jan Audun
title Drepanoistodus basiovalis
title_short Drepanoistodus basiovalis
title_full Drepanoistodus basiovalis
title_fullStr Drepanoistodus basiovalis
title_full_unstemmed Drepanoistodus basiovalis
title_sort drepanoistodus basiovalis
publisher Zenodo
publishDate 2021
url https://dx.doi.org/10.5281/zenodo.5558708
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geographic San Juan
Norway
Argentina
Rasmussen
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Italica
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genre Northern Sweden
genre_facet Northern Sweden
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spelling ftdatacite:10.5281/zenodo.5558708 2023-05-15T17:45:17+02:00 Drepanoistodus basiovalis Rasmussen, Jan Audun Eriksson, Mats E. Lindskog, Anders 2021 https://dx.doi.org/10.5281/zenodo.5558708 https://zenodo.org/record/5558708 unknown Zenodo http://zenodo.org/record/5557148 http://publication.plazi.org/id/FFB77B2CFFEEFFBF7D14FFF6FF91FFB7 http://zoobank.org/DE794E6F-E3E8-48A6-B42F-163DD2B675F8 https://zenodo.org/communities/biosyslit https://dx.doi.org/10.5852/ejt.2021.774.1533 http://zenodo.org/record/5557148 http://publication.plazi.org/id/FFB77B2CFFEEFFBF7D14FFF6FF91FFB7 https://dx.doi.org/10.5281/zenodo.5557160 https://dx.doi.org/10.5281/zenodo.5557154 https://dx.doi.org/10.5281/zenodo.5557158 http://zoobank.org/DE794E6F-E3E8-48A6-B42F-163DD2B675F8 https://dx.doi.org/10.5281/zenodo.5558707 https://zenodo.org/communities/biosyslit Open Access Creative Commons Zero v1.0 Universal https://creativecommons.org/publicdomain/zero/1.0/legalcode cc0-1.0 info:eu-repo/semantics/openAccess CC0 Biodiversity Taxonomy Animalia Chordata Conodonta Distacodontina Drepanoistodontidae Drepanoistodus Drepanoistodus basiovalis Text Taxonomic treatment article-journal ScholarlyArticle 2021 ftdatacite https://doi.org/10.5281/zenodo.5558708 https://doi.org/10.5852/ejt.2021.774.1533 https://doi.org/10.5281/zenodo.5557160 https://doi.org/10.5281/zenodo.5557154 https://doi.org/10.5281/zenodo.5557158 https://doi.org/10.5281/zenodo.5558707 2021-11-05T12:55:41Z Drepanoistodus basiovalis (Sergeeva, 1963) Fig. 6A–D Oistodus basiovalis Sergeeva, 1963: 96, pl. 7 figs 6–7, text-fig. 3. Drepanoistodus basiovalis – Lindström 1971: 43, text-figs 6, 8. — Stouge & Bagnoli 1990: 15, pl. 5 figs 18–24. — Dzik 1990: fig. 12; 1994: 78, pl. 16 figs. 16–20, text-fig. 12a; 2020: fig. 7A–E. — Rasmussen 1991: 277, fig. 6l; 2001: 71–73, pl. 5: 9 (cum. syn.). — Löfgren 1994: fig. 6.30; 2000a: fig. 4w; 2006: figs 3n, 3ab. — Viira et al . 2001: fig. 5z. — Zhen & Percival 2004: 93, fig. 11a–j. — Tolmacheva et al . 2013: pl. 3, fig. 24. partim Drepanoistodus basiovalis – Löfgren 1978: 55–56, pl. 1 figs 11–16 (only), non 17 (= D. contractus (Lindström, 1955)). — Olgun 1987: 49, pl. 6w (only). — Landing et al . 2003: fig. 4e (only). — Zhen et al . 2011: 222–227, fig. 12a?, b–n, p–q (only). cf. Drepanoistodus basiovalis – Zhang 1998: 61–62, pl. 5 figs 5–12 (unusually short upper margin of the base). ? Drepanoistodus basiovalis – Lehnert et al. 1998: 55, pl. 3 figs 6, 12 (12 may belong to Paroistodus originalis (Sergeeva, 1963)). — Boncheva et al . 2009: text-fig. 3.8 (broken element). — Albanesi & Ortega 2016: fig. 7(6) (shares characters with D. basiovalis and D . cf. balticus ). — Feltes et al . 2016: fig. 3ac. — Wu et al . 2018: fig. 5e (unusually long base compared to the cusp). non Drepanoistodus basiovalis – Gutiérrez-Marco et al . 2008: 153, figs 3.29–3.31 (may be Drepanoistodus cf. basiovalis or Drepanoistodus cf. suberectus (Branson & Mehl, 1933). — Hints et al . 2012: fig. 6h (= Drepanoistodus cf. suberectus ). — Wu et al . 2017: fig. 7u (= Drepanoistodus contractus (Lindström)). — Lindskog et al . 2020: fig. 7v–w (= Drepanoistodus viirae sp. nov.). Original diagnosis (translated from Sergeeva, 1963 [in Russian]) Inclined conodonts, almost symmetrical, with a wide shortened base, the edge of which is rounded. Material examined 33 geniculate elements including 24 from the Lynna section. Original description, slightly shortened (translated from Sergeeva, 1963 [in Russian]) Medium-sized conodonts (0.52–0.92 mm), inclined; the degree of inclination of the cusp is 45–60°, sometimes up to 80°. Base high, not very long, elongated along the CD; base length 2.5–3 times its height ( comment by the authors: “ we find the meaning of the latter measure ambiguous ”). Base wall slightly transparent near the edge, rounded. The angle between the sides AC is more than 90°; angle between AD 40–45°; corners are smoothly obtuse. Transverse in cross section, the base is oval, elongated along CD and compressed along L1L2. From the C side, the base is compressed, sometimes with a thin keel near the tip, with a small keel on side D. The sides of the base L1 and L2 are smooth and flat. Basal cavity is not always visible, it is wide, but not deep, without visible tops. The cusp is long, straight or slightly curved towards L1, sharply tapering towards the tip; compressed. The sides of the cusp are almost flat, with a welldeveloped longitudinal, wide carina on L1 and less developed carina on the side L2. The carinae usually run from the base to the tip of the cusp. Thin keels occur on the lower (D) and upper (C) parts of the cusp. Remarks In her original diagnosis, Sergeeva (1963) only included geniculate elements with a rounded basal margin in “ Oistodus ” basiovalis , which is also evident from the species epithet: basiovalis (meaning oval base). This interpretation of the geniculate element in Drepanoistodus basiovalis is followed here. Additional typical characters that may be added to the original species description include: anterior margin and upper anterior corner rounded or weakly rounded; cusp usually straight; a median or median to lower, longitudinal carina present on the inner (sometimes slightly concave) side of the element. Carina is more distinct in Darriwilian specimens than in Dapingian ones. Whereas angle A (Fig. 3) between the cusp and the upper margin is 29.6° with a standard deviation at 5.6, the mean ratio between the length of the free upper margin and free cusp (b/c ratio) reaches 0.40 with a standard deviation of 0.1 (Fig. 5B). Occurrence Drepanoistodus basiovalis occurs from the L. antivariabilis Zone (sample LY12-9) to the interzone (“uncertain interval”) between the L. variabilis Zone and the Y. crassus Zone sensu Lindskog et al. (2020) in the Lynna River section (sample LY12-21b; between LY12-21 and LY12-22). In addition, D. basiovalis has been reported from several other localities in Baltoscandia and Poland, and also outside the Baltica palaeocontinent, e.g., New Brunswick, Argentina, Australia and China (for references, see the synonymy list above). : Published as part of Rasmussen, Jan Audun, Eriksson, Mats E. & Lindskog, Anders, 2021, Middle Ordovician Drepanoistodus (Vertebrata, Conodonta) from Baltica, with description of three new species, pp. 106-134 in European Journal of Taxonomy 774 on pages 118-119, DOI: 10.5852/ejt.2021.774.1533, http://zenodo.org/record/5557148 : {"references": ["Sergeeva S. P. 1963. [Conodonts from the Lower Ordovician in the Leningrad region.] Paleontologicheshij Zhurnal 1963 (2): 93 - 108. [In Russian.]", "Lindstrom M. 1971. Lower Ordovician conodonts of Europe. Geological Society of America Memoirs 127: 21 - 61. https: // doi. org / 10.1130 / MEM 127 - p 21", "Stouge S. & Bagnoli G. 1990. Lower Ordovician (Volkhovian - Kundan) conodonts from Hagudden, northern Oland, Sweden. Palaeontographica Italica 77: 1 - 54.", "Dzik J. 1990. Conodont evolution in high latitudes of the Ordovician. Courier Forschungsinstitut Senckenberg 117: l - 28.", "Rasmussen J. A. 1991. Conodont stratigraphy of the Lower Ordovician Huk Formation at Slemmestad, southern Norway. Norsk Geologisk Tidsskrift 71: 265 - 88.", "Lofgren A. 1994. Arenig (Lower Ordovician) conodonts and biozonation in the eastern Siljan district, central Sweden. Journal of Paleontology 68: 1350 - 1368. https: // doi. org / 10.1017 / S 0022336000034338", "Viira V., Lofgren A., Magi S. & Wickstrom J. 2001. An Early to Middle Ordovician succession of conodont faunas at Maekalda, northern Estonia. Geological Magazine 138: 699 - 718. https: // doi. org / 10.1017 / S 0016756801005945", "Zhen Y. Y. & Percival I. G. 2004. Middle Ordovician (Darriwilian) conodonts from allochthonous limestones in the Oakdale Formation of central New South Wales. Alcheringa 28: 77 - 111. https: // doi. org / 10.1080 / 03115510408619276", "Tolmacheva T. Yu., Zaitsev A. V. & Alekseev A. S. 2013. Middle and Upper Ordovician conodonts of the Moscow Syneclise: new data on stratigraphy of the borehole Gavrilov Yam- 1 section. Stratigrafiya. Geologicheskaya Korrelyatsiya 21: 52 - 77. https: // doi. org / 10.1134 / S 0869593813040096", "Lofgren A. 1978. Arenigian and Llanvirnian conodonts from Jamtland, northern Sweden. Fossils and Strata 13: 1 - 129.", "Olgun O. 1987. Komponenten-Analyse und Conodonten-Stratigraphie der Orthoceratenkalksteine im Gebiet Falbygden, Vastergotland, Mittelschweden. Sveriges Geologiska Undersokning, Ser. Ca 70: 1 - 78.", "Landing E., Westrop S. R. & Kim D. H. 2003. First Middle Ordovician biota from southern New Brunswick: stratigraphic and tectonic implications for the evolution of the Avalon continent. Canadian Journal of Earth Sciences 40: 715 - 730. https: // doi. org / 10.1139 / E 03 - 009", "Zhen Y. Y., Wang Z. H., Zhang Y. D., Bergstrom S. M., Percival I. G. & Chen J. F. 2011. Middle to Late Ordovician (Darriwilian - Sandbian) conodonts from the Dawangou Section, Kalpin area of the Tarim Basin, northwestern China. Records of the Australian Museum 63: 203 - 266. https: // doi. org / 10.3853 / j. 0067 - 1975.63.2011.1586", "Zhang J. 1998. Conodonts from the Guniutan Formation (Llanvirnian) in Hubei and Hunan Provinces, south-central China. Stockholm Contributions in Geology 46: 1 - 161.", "Lehnert D., Keller M. & Bordonaro D. 1998. Early Ordovician conodonts from the southern Cuyania terrane (Mendoza Province, Argentina). In: H. Szaniawski (ed.) Proceedings of the Sixth European Conodont Symposium (ECOS VI). 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