Amiota (Phortica) Schiner 1862

Subgenus Phortica (sensu stricto) Schiner, 1862 Phortica Schiner, 1862: 433 (as a genus); Máca 2003: 251 (revised status). Type species: Drosophila variegata Fallén, 1823. Amiota ( Phortica ): Wheeler 1952: 167. Diagnosis. Additional plate present between cerci and tenth sternite; aedeagal median ro...

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Main Authors: Huang, Jia, Gong, Lu, Tsaur, Shun-Chern, Zhu, Lin, An, Keying, Chen, Hongwei
Format: Text
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Published: Zenodo 2019
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Online Access:https://dx.doi.org/10.5281/zenodo.5221215
https://zenodo.org/record/5221215
id ftdatacite:10.5281/zenodo.5221215
record_format openpolar
institution Open Polar
collection DataCite Metadata Store (German National Library of Science and Technology)
op_collection_id ftdatacite
language unknown
topic Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Diptera
Drosophilidae
Amiota
spellingShingle Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Diptera
Drosophilidae
Amiota
Huang, Jia
Gong, Lu
Tsaur, Shun-Chern
Zhu, Lin
An, Keying
Chen, Hongwei
Amiota (Phortica) Schiner 1862
topic_facet Biodiversity
Taxonomy
Animalia
Arthropoda
Insecta
Diptera
Drosophilidae
Amiota
description Subgenus Phortica (sensu stricto) Schiner, 1862 Phortica Schiner, 1862: 433 (as a genus); Máca 2003: 251 (revised status). Type species: Drosophila variegata Fallén, 1823. Amiota ( Phortica ): Wheeler 1952: 167. Diagnosis. Additional plate present between cerci and tenth sternite; aedeagal median rod developed, basally and medially with 1 or 2 pair(s) of bridges articulated with anterodorsal corners of gonopods (after Chen et al . 2007). ......continued on the next page Two or more putative morphospecies within a bracket pair indicate they (at least most of the samples) are delimited as a single species; only one putative morphospecies within a bracket pair indicates it is delimited as two or more species. Description. Male and female: Eye brownish red. Ocellar triangle dark brown to black. Frons pollinose, grayish brown to black, with a few interfrontal setulae medially. Fronto-orbital plate often silvery-white. Pedicel and first flagellomere grayish brown. Face grayish brown, with yellowish white patches on lower corners. Clypeus medially white to yellow, laterally dark brown to black. Gena grayish yellow to brown; postgena dark brown. Palpus somewhat triangular, brown basally, grayish yellow and with a few setae distally. Vibrissa prominent; other orals small. Mesoscutum and pleura grayish orange-brown, with brownish to black patches and pollinose patterns (Figs. 5, 6). Postpronotal lobe pale yellow, with 1 long and a few short setae. Acrostichal setulae in ca. 6–8 irregular rows. Prescutellar setae usually 1 pair. Scutellum usually concolorous with thorax, with dark brown to black patches (Figs. 5, 6). Basal scutellar setae divergent; apicals cruciate. Wing hyaline, sometimes smoky; veins grayish yellow. Basal medial-cubital crossvein present; C 1 setae 2, indistinctly differentiated. Costal vein with spinules on ventral surface between R 2+3 and R 4+5. R 2+3 slightly curved to costa at tip; R 4+5 distally convergent with M 1. Halters white. Legs yellow; femora usually brown to black except for apical portions; tibiae usually with 3 brown to black rings. Foreleg femur with 2 or 3 irregular rows of long setae on posterior surface. Preapical dorsal setae present on all tibiae. Midleg tarsus ventrally with 2 rows of minute cuneiform setulae on inner and outer sides; hindleg tarsus with 1 row of minute cuneiform setulae on underside; fore- and hindleg first tarsomeres each as long as three succeeding tarsomeres together; midleg first tarsomere as long as all tarsomeres together. Abdominal tergites yellow to orangeyellow (Figs. 5, 6): second to fifth tergites with broad brownish to black bands on posterior margins; sixth tergite nearly entirely dark, narrow anterolaterally. Sternites usually grayish yellow. Male terminalia (Figs. 7–37): Epandrium almost not constricted mid-dorsally, with pubescence and setae; apodeme developed along anterior margins. Cercus almost oval, separated from epandrium, entirely pubescent and setigerous. Surstylus with numerous setae on outer surface. Membrane between epandrium and cercus pubescent. Hypandrium arched, usually with 1 pair of apodeme processes on anterior portion, posteriorly contiguous to lateral corners of gonopods. Gonopods fused, forming posteromedian plate, anteriorly forming vertical process. Parameres usually basally contiguous to anterior portion of hypandrium and tips of distally bifurcated ventral branch of aedeagal apodeme. Aedeagus composed of outer membranous tube and more or less sclerotized median rod; outer membrane posteriorly connect to vertical process of gonopod; median rod basally and/or medially connect to basal corners of gonopods by 1 or 2 pair(s) of bridges (2 pairs in Figs. 12C, 20B, 20D, 30C, 33D, 36C, 37C) and basally contiguous to dorsal branch of aedeagal apodeme; basal bridge sometimes with sclerotized branch (Figs. 7E, 10C, 15E); ventral bridge (termed inner paraphysis in Bächli et al . 2004) usually contiguous to medial process of aedeagus (Figs. 30C, 33D, 37C), sometimes elongated and dilated apically (Figs. 12C, 20B, 20D, 36C). Female specimens were mostly collected from tree trunks, and were difficult to identify to species by morphological characters only (Chen & Toda 1997). Therefore, they are still rarely known in this study. Remarks. For the new species described below, only the characters inconsistent with the aforementioned general descriptions are provided for brevity. In the species descriptions, an asterisk (*) denotes a new record. : Published as part of Huang, Jia, Gong, Lu, Tsaur, Shun-Chern, Zhu, Lin, An, Keying & Chen, Hongwei, 2019, Revision of the subgenus Phortica (sensu stricto) (Diptera, Drosophilidae) from East Asia, with assessment of species delimitation using DNA barcodes, pp. 1-75 in Zootaxa 4678 (1) on pages 17-20, DOI: 10.11646/zootaxa.4678.1.1, http://zenodo.org/record/3468782 : {"references": ["Schiner, I. R. (1862) Vorlaufiger Commentar zum dipterologischen Theile der \" Fauna Austriaca \". Wiener Entomologische Monatschrift, 6, 428 - 436.", "Maca, J. (2003) Taxonomic notes on the genera previously classified in the genus Amiota Loew (Diptera, Drosophilidae, Steganinae). Acta Universitatis Carolinae Biologica, 47, 247 - 274.", "Fallen, J. C. (1823) Sveciae, Geomyzides. Litteris Berlingianis, Lundae, 8 pp.", "Wheeler, M. R. (1952) The Drosophilidae of the Nearctic Region, exclusive of the genus Drosophila. The University of Texas Publication, 5204, 162 - 218.", "Chen, H. W., Toda, M. J., Lakim, M. B. & Mohamed, M. B. (2007) The Phortica sensu stricto (Insecta: Diptera: Drosophilidae) from Malaysia. The Raffles Bulletin of Zoology, 55, 23 - 41.", "Bachli, G., Vilela, C. R., Escher, S. A. & Saura, A. (2004) The Drosophilidae (Diptera) of Fennoscandia and Denmark. In: Fauna Entomologica Scandinavica. Vol. 39. Brill, Leiden and New York, 362 pp.", "Chen, H. W. & Toda, M. J. (1997) Amiota (Phortica) magna species-complex, with descriptions of three new species from China (Diptera, Drosophilidae). Japanese Journal of Entomology, 65, 784 - 792."]}
format Text
author Huang, Jia
Gong, Lu
Tsaur, Shun-Chern
Zhu, Lin
An, Keying
Chen, Hongwei
author_facet Huang, Jia
Gong, Lu
Tsaur, Shun-Chern
Zhu, Lin
An, Keying
Chen, Hongwei
author_sort Huang, Jia
title Amiota (Phortica) Schiner 1862
title_short Amiota (Phortica) Schiner 1862
title_full Amiota (Phortica) Schiner 1862
title_fullStr Amiota (Phortica) Schiner 1862
title_full_unstemmed Amiota (Phortica) Schiner 1862
title_sort amiota (phortica) schiner 1862
publisher Zenodo
publishDate 2019
url https://dx.doi.org/10.5281/zenodo.5221215
https://zenodo.org/record/5221215
long_lat ENVELOPE(12.989,12.989,66.233,66.233)
geographic Saura
geographic_facet Saura
genre Fennoscandia
genre_facet Fennoscandia
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spelling ftdatacite:10.5281/zenodo.5221215 2023-05-15T16:12:27+02:00 Amiota (Phortica) Schiner 1862 Huang, Jia Gong, Lu Tsaur, Shun-Chern Zhu, Lin An, Keying Chen, Hongwei 2019 https://dx.doi.org/10.5281/zenodo.5221215 https://zenodo.org/record/5221215 unknown Zenodo http://zenodo.org/record/3468782 http://publication.plazi.org/id/FFB5FFB09A71675AFF87FFAAFFFAFFB6 http://zoobank.org/91B46647-B441-41F5-BE27-3A67D699788C https://zenodo.org/communities/biosyslit https://dx.doi.org/10.11646/zootaxa.4678.1.1 http://zenodo.org/record/3468782 http://publication.plazi.org/id/FFB5FFB09A71675AFF87FFAAFFFAFFB6 https://dx.doi.org/10.5281/zenodo.3468794 https://dx.doi.org/10.5281/zenodo.3468796 https://dx.doi.org/10.5281/zenodo.3468798 https://dx.doi.org/10.5281/zenodo.3468800 https://dx.doi.org/10.5281/zenodo.3468802 https://dx.doi.org/10.5281/zenodo.3468804 https://dx.doi.org/10.5281/zenodo.3468806 https://dx.doi.org/10.5281/zenodo.3468808 https://dx.doi.org/10.5281/zenodo.3468810 https://dx.doi.org/10.5281/zenodo.3468812 https://dx.doi.org/10.5281/zenodo.3468814 https://dx.doi.org/10.5281/zenodo.3468816 https://dx.doi.org/10.5281/zenodo.3468818 https://dx.doi.org/10.5281/zenodo.3468820 https://dx.doi.org/10.5281/zenodo.3468822 https://dx.doi.org/10.5281/zenodo.3468824 https://dx.doi.org/10.5281/zenodo.3468826 https://dx.doi.org/10.5281/zenodo.3468828 https://dx.doi.org/10.5281/zenodo.3468830 https://dx.doi.org/10.5281/zenodo.3468832 https://dx.doi.org/10.5281/zenodo.3468834 https://dx.doi.org/10.5281/zenodo.3468836 https://dx.doi.org/10.5281/zenodo.3468838 https://dx.doi.org/10.5281/zenodo.3468840 https://dx.doi.org/10.5281/zenodo.3468842 https://dx.doi.org/10.5281/zenodo.3468844 https://dx.doi.org/10.5281/zenodo.3468846 https://dx.doi.org/10.5281/zenodo.3468848 https://dx.doi.org/10.5281/zenodo.3468850 https://dx.doi.org/10.5281/zenodo.3468852 https://dx.doi.org/10.5281/zenodo.3468854 https://dx.doi.org/10.5281/zenodo.3468856 https://dx.doi.org/10.5281/zenodo.3468858 http://zoobank.org/91B46647-B441-41F5-BE27-3A67D699788C https://dx.doi.org/10.5281/zenodo.5221214 https://zenodo.org/communities/biosyslit Open Access info:eu-repo/semantics/openAccess Biodiversity Taxonomy Animalia Arthropoda Insecta Diptera Drosophilidae Amiota Text Taxonomic treatment article-journal ScholarlyArticle 2019 ftdatacite https://doi.org/10.5281/zenodo.5221215 https://doi.org/10.11646/zootaxa.4678.1.1 https://doi.org/10.5281/zenodo.3468794 https://doi.org/10.5281/zenodo.3468796 https://doi.org/10.5281/zenodo.3468798 https://doi.org/10.5281/zenodo.3468800 https: 2021-11-05T12:55:41Z Subgenus Phortica (sensu stricto) Schiner, 1862 Phortica Schiner, 1862: 433 (as a genus); Máca 2003: 251 (revised status). Type species: Drosophila variegata Fallén, 1823. Amiota ( Phortica ): Wheeler 1952: 167. Diagnosis. Additional plate present between cerci and tenth sternite; aedeagal median rod developed, basally and medially with 1 or 2 pair(s) of bridges articulated with anterodorsal corners of gonopods (after Chen et al . 2007). ......continued on the next page Two or more putative morphospecies within a bracket pair indicate they (at least most of the samples) are delimited as a single species; only one putative morphospecies within a bracket pair indicates it is delimited as two or more species. Description. Male and female: Eye brownish red. Ocellar triangle dark brown to black. Frons pollinose, grayish brown to black, with a few interfrontal setulae medially. Fronto-orbital plate often silvery-white. Pedicel and first flagellomere grayish brown. Face grayish brown, with yellowish white patches on lower corners. Clypeus medially white to yellow, laterally dark brown to black. Gena grayish yellow to brown; postgena dark brown. Palpus somewhat triangular, brown basally, grayish yellow and with a few setae distally. Vibrissa prominent; other orals small. Mesoscutum and pleura grayish orange-brown, with brownish to black patches and pollinose patterns (Figs. 5, 6). Postpronotal lobe pale yellow, with 1 long and a few short setae. Acrostichal setulae in ca. 6–8 irregular rows. Prescutellar setae usually 1 pair. Scutellum usually concolorous with thorax, with dark brown to black patches (Figs. 5, 6). Basal scutellar setae divergent; apicals cruciate. Wing hyaline, sometimes smoky; veins grayish yellow. Basal medial-cubital crossvein present; C 1 setae 2, indistinctly differentiated. Costal vein with spinules on ventral surface between R 2+3 and R 4+5. R 2+3 slightly curved to costa at tip; R 4+5 distally convergent with M 1. Halters white. Legs yellow; femora usually brown to black except for apical portions; tibiae usually with 3 brown to black rings. Foreleg femur with 2 or 3 irregular rows of long setae on posterior surface. Preapical dorsal setae present on all tibiae. Midleg tarsus ventrally with 2 rows of minute cuneiform setulae on inner and outer sides; hindleg tarsus with 1 row of minute cuneiform setulae on underside; fore- and hindleg first tarsomeres each as long as three succeeding tarsomeres together; midleg first tarsomere as long as all tarsomeres together. Abdominal tergites yellow to orangeyellow (Figs. 5, 6): second to fifth tergites with broad brownish to black bands on posterior margins; sixth tergite nearly entirely dark, narrow anterolaterally. Sternites usually grayish yellow. Male terminalia (Figs. 7–37): Epandrium almost not constricted mid-dorsally, with pubescence and setae; apodeme developed along anterior margins. Cercus almost oval, separated from epandrium, entirely pubescent and setigerous. Surstylus with numerous setae on outer surface. Membrane between epandrium and cercus pubescent. Hypandrium arched, usually with 1 pair of apodeme processes on anterior portion, posteriorly contiguous to lateral corners of gonopods. Gonopods fused, forming posteromedian plate, anteriorly forming vertical process. Parameres usually basally contiguous to anterior portion of hypandrium and tips of distally bifurcated ventral branch of aedeagal apodeme. Aedeagus composed of outer membranous tube and more or less sclerotized median rod; outer membrane posteriorly connect to vertical process of gonopod; median rod basally and/or medially connect to basal corners of gonopods by 1 or 2 pair(s) of bridges (2 pairs in Figs. 12C, 20B, 20D, 30C, 33D, 36C, 37C) and basally contiguous to dorsal branch of aedeagal apodeme; basal bridge sometimes with sclerotized branch (Figs. 7E, 10C, 15E); ventral bridge (termed inner paraphysis in Bächli et al . 2004) usually contiguous to medial process of aedeagus (Figs. 30C, 33D, 37C), sometimes elongated and dilated apically (Figs. 12C, 20B, 20D, 36C). Female specimens were mostly collected from tree trunks, and were difficult to identify to species by morphological characters only (Chen & Toda 1997). Therefore, they are still rarely known in this study. Remarks. For the new species described below, only the characters inconsistent with the aforementioned general descriptions are provided for brevity. In the species descriptions, an asterisk (*) denotes a new record. : Published as part of Huang, Jia, Gong, Lu, Tsaur, Shun-Chern, Zhu, Lin, An, Keying & Chen, Hongwei, 2019, Revision of the subgenus Phortica (sensu stricto) (Diptera, Drosophilidae) from East Asia, with assessment of species delimitation using DNA barcodes, pp. 1-75 in Zootaxa 4678 (1) on pages 17-20, DOI: 10.11646/zootaxa.4678.1.1, http://zenodo.org/record/3468782 : {"references": ["Schiner, I. R. (1862) Vorlaufiger Commentar zum dipterologischen Theile der \" Fauna Austriaca \". Wiener Entomologische Monatschrift, 6, 428 - 436.", "Maca, J. (2003) Taxonomic notes on the genera previously classified in the genus Amiota Loew (Diptera, Drosophilidae, Steganinae). Acta Universitatis Carolinae Biologica, 47, 247 - 274.", "Fallen, J. C. (1823) Sveciae, Geomyzides. Litteris Berlingianis, Lundae, 8 pp.", "Wheeler, M. R. (1952) The Drosophilidae of the Nearctic Region, exclusive of the genus Drosophila. The University of Texas Publication, 5204, 162 - 218.", "Chen, H. W., Toda, M. J., Lakim, M. B. & Mohamed, M. B. (2007) The Phortica sensu stricto (Insecta: Diptera: Drosophilidae) from Malaysia. The Raffles Bulletin of Zoology, 55, 23 - 41.", "Bachli, G., Vilela, C. R., Escher, S. A. & Saura, A. (2004) The Drosophilidae (Diptera) of Fennoscandia and Denmark. In: Fauna Entomologica Scandinavica. Vol. 39. Brill, Leiden and New York, 362 pp.", "Chen, H. W. & Toda, M. J. (1997) Amiota (Phortica) magna species-complex, with descriptions of three new species from China (Diptera, Drosophilidae). Japanese Journal of Entomology, 65, 784 - 792."]} Text Fennoscandia DataCite Metadata Store (German National Library of Science and Technology) Saura ENVELOPE(12.989,12.989,66.233,66.233)