Canis aureus Linnaeus 1758
Canis aureus Linnaeus, 1758 Golden Jackal Canis aureus Linnaeus, 1758:40. Type locality “oriente;” restricted to “Benná Mts., Laristan, S. Persia ” by Thomas (1911:135). Lupus aureus Linnaeus, 1758:40. No type locality. From a pre- 1758 name. Vulpes indiae orientalis Linnaeus, 1758:41. No type local...
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Biodiversity Taxonomy Animalia Chordata Mammalia Carnivora Canidae Canis Canis aureus |
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Biodiversity Taxonomy Animalia Chordata Mammalia Carnivora Canidae Canis Canis aureus Moehlman, Patricia D. Hayssen, Virginia Canis aureus Linnaeus 1758 |
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Biodiversity Taxonomy Animalia Chordata Mammalia Carnivora Canidae Canis Canis aureus |
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Canis aureus Linnaeus, 1758 Golden Jackal Canis aureus Linnaeus, 1758:40. Type locality “oriente;” restricted to “Benná Mts., Laristan, S. Persia ” by Thomas (1911:135). Lupus aureus Linnaeus, 1758:40. No type locality. From a pre- 1758 name. Vulpes indiae orientalis Linnaeus, 1758:41. No type locality. From a pre-1758 name. [ Canis ] anthus F. Cuvier, 1820:plate. Type locality “ Senegal.” Canis variegatus Cretzschmar, 1826:31, plate 10. Type locality “Nubien. Oberes Egypten;” preoccupied. Canis syriacus Hemprich and Ehrenberg, 1830:sig z, plate 16. Type locality “e monte Lebano.” Canis lupaster Hemprich and Ehrenberg, 1833:sig ff. Type locality “Fayum, Egypt ” (not seen cited in Allen 1939:194). Canis sacer Hemprich and Ehrenberg, 1833:sig ff. Type locality “Fayum, Egypt ” (not seen cited in Allen 1939:194). Canis riparius Hemprich and Ehrenberg, 1833:sig ff. Type local- ity “Coast of Abyssinia, near Arkiko” (not seen cited in Allen 1939:195). Canis aureus indicus Hodgson, 1833:237. Type locality “Himalaya.” Canis aureus var. moreotica I. Geoffroy Saint-Hilaire, 1835:plate 1. Type locality “de Morée.” Thous anthus : Hamilton Smith, 1839:195. Name combination. Thous variegatus : Hamilton Smith, 1839:198. Name combination. Thous sengalensi Hamilton Smith, 1839:201, plate 13. Type locality “ Gambia and Senegal.” Sacalius aureus : Hamilton Smith, 1839:214. Name combination. Sacalius barbarus Hamilton Smith, 1839:218. Type locality “ Tunis;” preoccupied by Canis barbarus Shaw, 1800:311 synonym for Vulpes vulpes. Sacalius indicus : Hamilton Smith, 1839:219. Name combination. C [ anis ]. dalmatinus Wagner, 1841:383. No type locality; Dalmatia implied. Canis graecus Wagner, 1841:383. Type locality “Europaei.” Renaming of C. aureus moreotica I. Geoffroy Saint-Hilaire, 1835. C [ anis ]. aureus vulgaris Wagner, 1841:383. No type locality. Canis aureus algirensis Wagner, 1841:384. Type locality “d’Alger” from ‘chacal d’Alger’ I. Geoffroy Saint-Hilaire, 1835:22. C [ anis ]. aureus tripolitanus Wagner, 1841:384. No type locality; Tripoli, Tunis implied. Canis aureus typicus Kolenati, 1858:96. No type locality; Persia, Georgia, Turkey implied. Canis aureus typicus var caucasica Kolenati, 1858:96. No type locality; Caucasus implied. Canis aureus var syriaco Kolenati, 1858:96. Unauthorized emendation of syriacus Hemprich and Ehrenberg, 1830. Canis aureus var indica Kolenati, 1858:96. Unauthorized emendation of indicus Hodgson, 1833. Canis aureus var nubica Kolenati, 1858:96. Renaming of Canis variegatus Cretzschmar, 1826 but Kolenati attributes the name to Rüppell without giving a date. Canis aureus var algira Kolenati, 1858:96. Unauthorized emendation of algirensis Wagner, 1841. Canis aureus var senegalensis : Kolenati, 1858:96. Name combination. Canis aureus var barbarus : Kolenati, 1858:96. Name combination. Canis aureus balcanicus Brusina, 1892:317. Type locality “Nardu — nadomak Valpova — na obali Drave, Slavonija.” Canis hadramaticus Noack, 1896:356. Type locality “Hadramaut in Arabien.” A jackal-wolf composite. The jackal was chosen as the lectotype (Morrison-Scott 1939) Canis mengesi Noack, 1897:518. Type locality “Ostafrikanische.” Canis lupus minor Mojsisovico, 1897:202, 241, 244. Preoccupied by Canis spelaeus minor Wagner, 1829 according to Kretzoi, 1947. Canis cruesemanni Matschie, 1900:145. Type locality “ Siam.” Canis anthus soudanicus Thomas, 1903:295. Type locality “El Obeid,” Kordofan, Sudan. Canis somalicus Lorenz, 1906:306. Type locality “Ireso bei Agada.” Canis gallaënsis Lorenz, 1906:307. Type locality “Ginea (Arussi).” Canis doederleini Hilzheimer, 1906a:116. Type locality “Oberägypten.” Canis thooides Hilzheimer, 1906b:364. Type locality “Sennaar.” Canis lupaster grayi Hilzheimer, 1906b:367. Type locality “Nordafrika.” Canis studeri Hilzheimer, 1906b:368. Type locality “ Tunis.” C [ anis ]. mengesi lamperti Hilzheimer, 1906b:371. No type locality. Thos aureus bea Heller, 1914:5. Type locality “Loita Plains, British East Africa,” Kenya. Canis indicus kola Wroughton, 1916:651. Type locality “Palanpur,” India. Canis naria Wroughton, 1916:651. Type locality “Coorg,” west coast of India. Canis lanka Wroughton, 1916:652. Type locality “Mankeni,” Sri Lanka. Thos lupaster maroccanus Cabrera, 1921:263. Type locality “ Mogador (Marruecos).” Th [ os ]. l [ upaster ]. Algirensis : Cabrera, 1921:263. Name combination. Th [ os ]. a [ ureus ]. Riparius : Cabrera, 1921:264. Name combination. Th [ os ]. aureus nubianus Cabrera, 1921:264. Renaming of variegatus Cretzschmar, 1826. Canis aureus hungaricus Éhik, 1938:11. Type locality “Tyukod (Marshy-land Ecsed), Comitat Szatmár, Hungary;” Preoccupied according to Kretzoi, 1947. Canis aureus minor : Éhik, 1938:13. Name combination; preoccupied. Thos aureus algirensis : Allen, 1939:194. Name combination. Thos aureus anthus : Allen, 1939:194. Name combination. Thos aureus lupaster : Allen, 1939:194. Name combination. Thos aureus maroccanus : Allen, 1939:195. Name combination. Thos aureus soudanicus : Allen, 1939:195. Name combination. Thos gallaensis : Allen, 1939:195. Name combination. Thos lamperti : Allen, 1939:195. Name combination. [ Canis aureus ] moreoticus Kretzoi, 1947:287. Unjustified emendation of moreotica I. Geoffroy Saint-Hilaire, 1835. Thos aureus ecsedensis Kretzoi, 1947:287. Renaming of C. a. hungaricus Éhik, 1938. Canis lupaster doederleini : Saleh and Basuony, 2014:49. Name combination. Canis aureus qattarensis Saleh and Basuony, 2014:49. Type locality “northern region of the Egyptian Western Desert.” CONTENT AND CONTEXT. Order Carnivora, family Canidae, subfamily Caninae, genus Canis . As many as 13 subspecies of C . aureus are distinguished through Europe and Africa (Allen 1939; Ellerman and Morrison-Scott 1951; Coetzee 1977). However, these subspecies may be part of a 3-species complex (see “Nomenclatural Notes”). Unfortunately, much of the literature does not specify geographic location or subspecies. Here, C . aureus is used for the global range from Africa into Eurasia (Wozencraft 2005). Therefore, we list the historical 13 subspecies of C. aureus (Heller 1914; Allen 1939; Ellerman and Morrison- Scott 1951; Coetzee 1977; Wozencraft 2005). The newly named subspecies, qattarensis Saleh and Basuony, 2014, is tentatively treated as a synonym of syriacus . Subspecies designations for the orientalis Linnaeus, 1758 and studeri Hilzheimer, 1906b are not known. C. a. algirensis Wagner, 1841. See above; synonyms are algira Kolenati, 1858; barbarus (Hamilton Smith, 1839); grayi Hilzheimer, 1906b; tripolitanus Wagner, 1841. C. a. anthus F. Cuvier, 1820. See above; synonym is senegalensis (Hamilton Smith, 1839). C. a. aureus Linnaeus, 1758. See above; synonyms are balcanicus Brusina, 1892; caucasica Kolenati, 1858; hadramauticus Noack, 1896; maroccanus (Cabrera, 1921); orientalis (Linnaeus, 1758); typicus Kolenati, 1858; vulgaris Wagner, 1841. C. a. bea (Heller, 1914). See above. C. a. cruesemanni Matschie, 1900. See above. C. a. ecsedensis (Kretzoi, 1947). See above; synonyms are hungaricus Ehik, 1938; minor Mojsisovico, 1897 (preoccupied). C. a. indicus Hodgson, 1833. See above; synonyms are indica Kolenati, 1858; kola Wroughton, 1916. C. a. lupaster Hemprich and Ehrenberg, 1833. See above; synonym is sacer Hemprich and Ehrenberg, 1833. C. a. moreotica I. Geoffroy Saint-Hilaire, 1835. See above; synonyms are dalmatinus Wagner, 1841; graecus Wagner, 1841. C. a. naria Wroughton, 1916. See above; synonym is lanka Wroughton, 1916. C. a. riparius Hemprich and Ehrenberg, 1833. See above; synonyms are gallaënsis Lorenz, 1906; hagenbecki Noack, 1897; lamperti Hilzheimer, 1906b; mengesi Noack, 1897; somalicus Lorenz, 1906. C. a. soudanicus Thomas, 1903. See above, synonyms are doederleini Hilzheimer, 1906a; nubianus (Cabrera, 1921); nubica Kolenati, 1858; thooides Hilzheimer, 1906b; variegatus Cretzschmar, 1826 (preoccupied). C. a. syriacus Hemprich and Ehrenberg, 1830. See above, synonym are qattarensis Saleh and Basuony, 2014; syriaco Kolenati, 1858. NOMENCLATURAL NOTES. The above-named list may represent 3 taxa of canids: Canis anthus (African wolf), C. aureus , and C. lupus (gray wolf). Genetic analyses of C . aureus and C. lupus suggest that C . aureus from Africa and Eurasia are distinct clades and should be considered to be separate species, that is C. anthus (Africa) and C. aureus (Eurasia—Koepfli et al. 2015). As noted above, the literature does not distinguish these species and we include both as C. aureus . In addition, large C . aureus in Egypt ( lupaster ) may be a cryptic subspecies of C . anthus (Rueness et al. 2011; Gaubert et al. 2012). Canis aureus lupaster (Hemprich and Ehrenberg 1833) is classified as a golden jackal; however Ferguson (1981) suggested that the taxon C. aureus lupaster , which is present in arid areas of Egypt and Libya (Osborn and Helmy 1980), may actually represent a small C. lupus rather than a large jackal. Mitochondrial DNA analyses suggest that C. a. lupaster represents an ancient wolf lineage that may have colonized Africa prior to the northern hemisphere radiation (Rueness et al. 2011). Mitochondrial DNA analyses assigned individuals from Algeria, Mali, and Senegal to C . lupus lupaster (Gaubert et al. 2012) and identified C. lupus lupaster mtDNA haplotypes in C. aureus from Senegal questioning the genetic differentiation between the proposed C . anthus and C . aureus . We retain C. aureus lupaster as a form of the golden jackal not as a subspecies of C. lupus following Wozencraft (2005— Qumsiyeh 1996; Ferguson 2002). The specific name is Latin for golden. Common names include Asiatic jackal, common jackal, ibn awa or ibn awee (classic Arab), wa wie (spoken Arab), cakalli (Albanian), čagalj (Croatian), šakal obecný (Czech), sjakal (Danish, Swedish), jakhal (Dutch), šaakal (Estonian), sakaali (Finnish), Sjakalur (Faeroese), chacal doré (French), Goldschakal (German), tơɑκɑλ (Greek), aranysakál (Hungarian), sciacallo dorato (Italian), chagal, turg (Kurdish), zeltainais šakālis (Latvian), xakali (Maltese), gullsjakal (Nowegian), shoghal (Persian), szakal zlocisty (Polish), chacal-dourado (Portuguese), şakal (Rumanian), šakal obyčajný (Slovakian, Slovenian), chacal (Spanish), çakal (Turkish—Hatt 1959; P. D. Moehlman, in litt.). Indigenous names include Amharic: tera kebero (Ethiopia); Fulani: sundu; Hausa: dila; Hindi: Giddhad; Kanada: nuree; Kiswahili: bweha wa mbugani, bweha dhahabu (Tanzania); Marathi (India): kolha; Nepali (Nepal), Bengali, Gujarati and Kutchi (India): shiyal; Singhelese: nariya; Songhai: nzongo; Tamil (India): peria naree; Wolof: Tili (P. D. Moehlman, in litt.). DIAGNOSIS Canis aureus occurs sympatrically with the silver-backed jackal ( Canis mesomelas ) and the side-striped jackal ( Canis adustus ) in parts of East Africa. All have skull lengths of 141– 147 mm with minimal divergence in size (van Valkenburgh and Wayne 1994). C . mesomelas is distinguished by reddish flanks and limbs, and the skull may have a less developed parietal crest (Clutton-Brock et al. 1976), the angle of its ears which are more upright, and, usually, the prominent dark “silver” saddle composed of black and white hairs (Walton and Joly 2003). C . aureus may have a dark saddle seasonally, which is not prominent. C . adustus characteristically has shorter ears set more to the side of the skull, a pale side-stripe, and a white-tipped tail (Moehlman and Jhala 2013). C. lupus is much larger with a heavier body and variable pelage coloring. The skull is much larger with a welldeveloped interparietal crest, strong jaw, and large canine and carnassial teeth (Clutton-Brock et al. 1976). GENERAL CHARACTERS Canis aureus (Fig. 1) is a medium-sized canid, considered the most typical representative of the genus Canis (Clutton- Brock et al. 1976). Basic coat color is golden but varies from pale creamy yellow to a dark tawny hue on a seasonal basis. The pelage on the back is often a mixture of black, brown, and white hairs, such that they can appear to have a dark saddle similar to the C. mesomelas (Walton and Joly 2003; Moehlman and Jhala 2013). The long contour hairs have an extensive black tip and a wider penultimate white band (Pocock 1938). In winter, a copious under wool is present (Pocock 1938). C . aureus inhabiting rocky, mountainous terrain may have a grayer coat shade (Sheldon 1992). The belly and underparts are a lighter pale-ginger to cream. The tail is bushy with a tan to black tip. Females have 7–8 mammae (Poché et al. 1987), comprising 4 pairs (Sheldon 1992). Eye color varies from pale yellow to amber (Lewis et al. 1968). Means and parenthetical ranges of external measurements (mm) from 6 males and 3 females, respectively, in Gujarat, India (Y. V. Jhala, in litt.) were: length of head and body, 793 (760–840), 760 (740–800); length of tail, 220 (200–240), 205 (200–210); length of ear, 76 (68–90), 80 (75–85); body mass (kg) of 6 male and 4 females was 8.8 (7.6–9.8) and 7.3 (6.5– 7.8), respectively. In Tanzania, the external measurements (mm or kg) from 2 males in the Serengeti (Moehlman and Jhala 2013) were: length of head and body, 740, 785; length of tail, 270, 280; length of ear, 110, 110; mass, 6.3, 7.7. In Egypt, means and parenthetical ranges (mm) for 9 adult C. a. lupaster of unknown sex were: length of head and body, 872 (822–893); length of tail, 312 (290–347); length of hind foot, 200 (190–212); length of ear, 112 (104–121); and body mass (kg) of 4 adults was 13 (10–15—Osborn and Helmy 1980). In Lebanon, external measurements (mm) of 3 adult C. a. syriacus were: total length, 988.3 (950–1015); length of tail, 275 (265–290); length of head and body, 713.3 (680–750); length of hind foot, 156 (140–170); length of ear from notch, 74.7 (70–80— Lewis et al. 1968). In Bangladesh, mean measurements (mm, SD , n ) of females and males, respectively, were: total length, 1009.3 (33.67, 6), 1058.6 (20.30, 7); length of tail, 220.1 (17.16, 7), 245.0 (21.41, 6); length of hind paw, 58.0 (4.93, 7), 56.3 (4.83, 7); length of hind foot, 146.2 (24.66, 5), 162.6 (8.52, 7); length of ear, 69.4 (4.80, 5), 74.1 (5.49, 7); mass (kg), 8.5 (0.73, 7), 10.3 (0.95, 7— Poché et al. 1987). Mean adult mass (kg) of 5 females and 3 males, respectively, was 10.0 and 12.0 (Palmqvist et al. 2002). On average, females are 12% lighter than males: 5.8 kg versus 6.6 kg (Moehlman and Hofer 1997). Mean cranial measurements (mm, SD Fig. 2) on 10 mandibles and upper jaws of adult C . aureus in Iran were: length of skull, 169, 31.9; length of cranium, 101, 23.7; nasal length, 67, 16.6; cranial width, 72, 4.4; length of mandible, 112, 34.6; lateral alveolar root to mental foramen, 25, 1.7; mental foramen to caudal mandibular border, 87, 3.8; mandibular foramen to base of mandible, 12, 0.4; caudal border of mandible to below mandibular foramen, 11, 0.8; maximum mandibular height, 60, 5.8 (Mofared 2013). Mean measurements (mm, range) for 3 Lebanese adults were: greatest length of skull, 148.6 (135.2–157.4); condylobasal length, 142.9 (131.9–149.1); zygomatic breadth, 83.9 (73.3–90.2); breadth of braincase, 48.8 (46.0–50.3); interorbital constriction, 29.4 (25.5–32.7); maxillary toothrow, 62.9 (60.4– 65.0); mandibular toothrow, 72.2 (58.2–74.9); mandibular length, 114.1 (104.1–121.0— Lewis et al. 1968). Mean cranial measurements (mm; with range and n ) of Egyptian adult C. a. lupaster were: condyloincisive length, 185.2 (173.5–196.0, 13); zygomatic width, 101.4 (93.5–111.3, 14); rostral width, 33.8 (31.2–37.8, 14); postorbital width, 34.8 (31.8–38.9, 13); basioccipital width at tempoparietal suture, 54.4 (51.8–59.1, 13); nasal length, 72.0 (65.9–84.8, 14); width across upper molars, 58.2 (54.2–63.0, 14); distance from anterior most surface of canine to posterior of 2nd molar, 80.3 (75.6–86.8, 14); skull height, 66.8 (62.0–74.0, 9—Osborn and Helmy 1980). In Bulgaria, mean cranial and tooth measurements (mm) for 92 males and 56 females, respectively, were: total skull length, 166, 161; neurocranium length, 89, 86.5; greatest length of nasals, 59.7, 57.9; greatest neurocranium breath, 52.5, 52.6; total length of mandible, 121.9, 118.6; height of vertebral ramus, 48.6, 46.7; length of P1 to P4, 33.9, 33.4; length of M1 to M3, 31.8, 31.0 (Stoyanev 2012). Mean mandible length and depth at 2nd molar (mm) for 5 females and 3 males, respectively, was f: 102.8, 14.3; m: 108.9, 15.3 (Palmqvist 2002). Condylobasal length (mm) from 10 males was 154, whereas it was 140 from 7+ female skulls (Pocock 1938). DISTRIBUTION Canis aureus is widespread in North and East Africa, Europe, the Middle East, Central Asia, and Southeast Asia (Fig. 3). They range from sea level in Eritrea to 3,500 m in the Bale Mo : Published as part of Moehlman, Patricia D. & Hayssen, Virginia, 2018, Canis aureus (Carnivore: Canidae), pp. 14-25 in Mammalian Species 50 (957) on pages 14-22, DOI: 10.1093/mspecies/sey002, http://zenodo.org/record/4573460 : {"references": ["LINNAEUS, C. 1758. Systema naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Editio decima, reformata. Volume 1. Laurentii Salvii, Stockholm, Sweden.", "CUVIER, F. 1820. Le chacal du Senegal, femelle. Pp. 1 - 3 in Histoire naturelle des mammiferes, avec des figures dessinees d'apres les animaux vivants, tome duexieme (E. Geoffroy Saint-Hilaire and F. Cuvier, eds.). A. Belin, Paris, France. 1824 on title page but material on the jackal is June 1820.", "CRETZSCHMAR, P. J. 1826. Atlas zu der Reise im nordlichen Afrika von Eduard Ruppell. Erste Abtheilung Zoologie. Gedruckt und in Commission bei Heinr. Ludw. Bronner, Frankfurt, Germany.", "ALLEN, G. M. 1939. A checklist of African mammals. Bulletin of the Museum of Comparative Zoology at Harvard College 83: 1 - 763.", "HODGSON, B. H. 1833. Description of the wild dog of the Himalaya. Asiatic Research 18: 221 - 237.", "HAMILTON SMITH, C. 1839. The natural history of dogs: Canidae or genus Canis of authors; including also the genera Hyaena and Proteles. Pp. 77 - 267 in Jardine's naturalist library. Volume 18. Mammalia. Dogs. Volume 1 (W. Jardine, ed.). W. H. Lizars, Edinburgh, Scotland.", "SHAW, G. 1800. General zoology or systematic natural history. Volume 1, part 2. G. Kearsley, London, United Kingdom.", "KOLENATI, F. A. 1858. Reiseerinnerungen. Die Bereisung Hocharmeniens und Elisabethopols, der Schekinschen Provinz und des Kasbek im Central-Kaukasus. Verlagsbuchhandlung von Rudolf Kuntze, Dresden, Germany.", "BRUSINA, S. 1892. Cagalj balkanski (Canis aureus balcanicus Brus n. form.) iz Slavonije. Glasnik Hrvatskoga Naravoslovnoga Drustva Zagreb 7: 316 - 317.", "NOACK, T. 1896. I. Wissenschaftliche Mittheilungen. 1. Ein neuer Steinbock und ein neuer (?) [sic] Canide aus Arabien. Zoologischer Anzeiger 19: 353 - 356.", "NOACK, T. 1897. Ostafrikanische Schakale. Zoologischer Anzeiger 20: 517 - 521.", "MOJSISOVICO, A. 1897. Das Thierleben der Osterreichisch-Ungarischen Tiefebenen. Biologische und thiergeographische Skizzen und Bilder. A. Holder, Wien, Austria.", "KRETZOI, M. 1947. New names for mammals (Notes on nomenclature No. 3). Annales Historico-Naturales Musei Nationalis Hungarici 40: 285 - 287.", "MATSCHIE, P. 1900. Herr Matschie sprach uber den Schakal des Menam-Gebietes in Siam. Sitzungs-Berichte der Gesellschaft der Naturforschender Freunde zu Berlin 3: 144 - 145.", "LORENZ, L. 1906. Uber zwei neue Schakale aus Nordostafrika aus der Sammlung des Freiherrn C. von Erlanger. Anzeiger, Kaiserliche Akademie der Wissenschaften in Wien. Mathematisch- Naturwissenschaftliche Klasse 43: 306 - 310.", "HILZHEIMER, M. 1906 a. Papio mundamensis, Felis deliensis, Canis reissii und andre neue Saugetiere. Zoologischer Anzeiger 30: 109 - 118.", "HILZHEIMER, M. 1906 b. Die geographische Verbreitung der Afrikanischen Grauschakale. Zoologischer Beobachter 47: 363 - 373.", "HELLER, E. 1914. New subspecies of mammals from equatorial Africa. Smithsonian Miscellaneous Collections 63: 1 - 12.", "CABRERA, A. 1921. Algunos carnivoros africanos nuevos. Boletin de la Real Sociedad Espanola de Historia Natural, Madrid 21: 261 - 264.", "EHIK, J. 1938. Sakal vagy nadifarkas hazankbol [Jackal or reed-wolf from Hungary]. Annales Musei Nationalis Hungarici, Pars Zoological 31: 11 - 15.", "COETZEE, C. G. 1977. Order Carnivora. Part 8. Pp. 1 - 42 in The mammals of Africa: an identification manual (J. Meester and H. W Setzer, eds.). Smithsonian Institution Press, Washington, D. C.", "RUENESS, E. K., ET AL. 2011. The cryptic African wolf: Canis aureus lupaster is not a golden jackal and is not endemic to Egypt. PLoS One 6: e 16385.", "GAUBERT, P., ET AL. 2012. Reviving the African wolf Canis lupus lupaster in North and West Africa: a mitochondrial lineage ranging more than 6,000 km wide. PLoS One 7: e 42740.", "FERGUSON, W. W. 1981. The systematic position of Canis aureus lupaster (Carnivora: Canidae) and the occurrence of Canis lupus in North Africa, Egypt and Sinai. Mammalia 45: 459 - 465.", "QUMSIYEH, M. B. 1996. Mammals of the Holy Land. Texas Tech University Press, Lubbock.", "FERGUSON, W. W. 2002. The mammals of Israel. Gefen Publishing, Jeruslaem, Israel.", "CLUTTON- BROCK, J., G. B. CORBET, AND M. HILLS. 1976. A review of the family Canidae, with a classification by numerical methods. Bulletin of the British Museum (Natural History) Zoology 29: 1 - 199.", "POCOCK, R. I. 1938. The jackals of S. W. Asia and S. E. Europe. Proceedings of the Zoological Society of London 108: 37 - 40.", "SHELDON, J. W. 1992. Wild dogs: the natural history of the non-domestic Canidae. Academic Press, Chicago, Illinois.", "POCHE, R. M., S. J. EVANS, P. SULTANA, M. E. HAGUE, R. STERNER, AND M. A. SIDDIQUE. 1987. Notes on the golden jackal (Canis aureus) in Bangladesh. Mammalia 51: 259 - 270.", "LEWIS, R. E., J. H. LEWIS, AND S. I. ATALLAH. 1968. A review of Lebanese mammals. Carnivora, Pinnipedia, Hyracoidea and Artiodactyla. Journal of Zoology (London) 154: 517 - 531.", "PALMQVIST, P., M. MENDOZA, A. ARRIBAS, AND D. R. GRoCKE. 2002. Estimating the body mass of Pleistocene canids: discussion of some methodological problems and a new ' taxon free' approach. Lethaia 35: 358 - 360.", "MOFARED, A. L. 2013. Macro-anatomical investigation of the skull of golden jackal (Canis aureus) and its clinical application during regional anesthesia. Global Veterinaria 10: 547 - 550.", "STOYANEV, S. 2012. Craniometric differentiation of golden jackals (Canis aureus L., 1758) in Bulgaria. Pp. 39 - 47 in International symposium on hunting, modern aspects of sustainable management of game populations, Zemun-Belgrade, Serbia.", "PRATER, S. 1971. The book of Indian animals. Bombay Natural History Society, Bombay, India.", "ADMASU, E., S. J. THIRGOOD, A. BEKELE, AND K. M. LAURENSON. 2004. Spatial ecology of golden jackal in farmland in the Ethiopian highlands. African Journal of Ecology 42: 144 - 152.", "MALEZ, V. 1984. The zooarcheological data as the base of colonizing the Markova cave on the island of Hvar. Pp. 617 - 621 in 9 th Yugoslavian speleological congress proceedings, Yugoslavia (in Croatian).", "RUTKOWSKI, R., ET AL. 2015. A European concern? Genetic structure and expansion of golden jackals (Canis aureus) in Europe and the Caucasus. PLoS One 10: e 0141236.", "FABBRI, E., ET AL. 2014. Genetic structure and expansion of golden jackals (Canis aureus) in the north-western distribution range (Croatia and eastern Italian Alps). Conservation Genetics 15: 187 - 199.", "ARNOLD, J., A. HUMER, M. HELTAI, D. MURARIU, N. SPASSOV, AND K. HaCKLANDER. 2012. Current status and distribution of golden jackals Canis aureus in Europe. Mammal Review 42: 1 - 11.", "KRYsTUFEK, B., D. MURARIU, AND C. KURTONUR. 1997. Present distribution of the golden jackal Canis aureus in the Balkans and adjacent regions. Mammal Review 27: 109 - 114.", "HOOIJER, D. A. 1961. The fossil vertebrates of Ksar'Akil, a Paleolithic rockshelter in Lebanon. Zoologische Verhandelingen 49: 1 - 68.", "SAVAGE, R. J. 1977. Evolution in carnivorous mammals. Paleontology 20: 237 - 271.", "KURTEN, B. 1974. A history of coyote-like dogs (Canidae, Mammalia). Acta Zoologica Fennica 140: 1 - 38.", "SPASSOV, N. 1989. The position of jackals in the Canis genus and lifehistory of the golden jackal (Canis aureus L.) in Bulgaria and the Balkans. Historia Naturalis Bulgarica, Sofia 1: 44 - 56.", "EWER, R. F. 1956. The fossil carnivores of the Transvaal caves: Canidae. Proceedings of the Zoological Society of London 126: 97 - 120.", "DIXSON, A. F. 1995. Baculum length and copulatory behaviour in carnivores and pinnipeds (Grand Order Ferae). Journal of Zoology (London) 235: 67 - 76.", "FULLER, T. K., A. R. BIKNEVICIUS, P. W. KAT, B. VAN VALKENBURGH, AND R. K. WAYNE. 1989. The ecology of three sympatric jackal species in the Rift Valley of Kenya. African Journal of Ecology 27: 313 - 323.", "SEITZ, V. A. 1959. Beobachtungen an Handaufgezogenen Goldschakalen (Canis aureus aligirensis). Zeitschrift fur Tierpsychologie 16: 747 - 771.", "MOEHLMAN, P. D. 1983. Socioecology of silverbacked and golden jackals (Canis mesomelas and Canis aureus). Pp. 423 - 453 in Advances in the study of mammalian behavior (J. F. Eisenberg and D. G. Kleiman, eds.). American Society of Mammalogists Special Publications, Lawrence, Kansas.", "MOEHLMAN, P. D. 1986. Ecology of cooperation in canids. Pp. 64 - 86 in Ecological aspects of social evolution (D. I. Rubenstein and R. W. Wrangham, eds.). Princeton University Press, Princeton, New Jersey.", "RAICHEV, E. 2011. Effect of shooting on the structure of population of golden jackal (Canis aureus L.) in Sarnena Sredna Gora Mountain. Agricultural Science and Technology 3: 276 - 280.", "FLOWER, S. S. 1932. Notes on the recent mammals of Egypt, with a list of the species recorded from that kingdom. Proceedings of the Zoological Society of London 1932: 369 - 450.", "Ginsberg, J. R., AND D. W. Macdonald (eds.). 1990. Foxes, wolves, jackals, and dogs. An action plan for the conservation of canids. International Union for the Conservation of Nature, Gland, Switzerland.", "MOEHLMAN, P. D. 1989. Intraspecific variation in canid social systems. Pp. 143 - 163 in Carnivore behavior, ecology, and evolution (J. L. Gittleman, ed.). Chapman & Hall, London, United Kingdom.", "KROFEL, M. 2008. Survey of golden jackals (Canis aureus L.) in northern Dalmatia, Croatia: preliminary results. Natura Croatia 17: 259 - 264.", "KROFEL, M. 2007. Golden jackals (Canis aureus L.) on the Peljesac Peninsula (southern Dalmatia, Croatia). Natura Croatica 16: 201 - 204.", "KHIDAS, K. 1990. Contribution a la connaissance du chacal dore. Facteurs modulant l'organisation sociale et territorial de la sous-espece algerienne (Canis aureus algirensis Wagner, 1841). Mammalia 54: 361 - 375.", "JAEGER, M. M., R. K. PANDIT, AND E. HAQUE. 1996. Season differences in territorial behavior by golden jackals in Bangladesh: howling versus confrontation. Journal of Mammalogy 77: 768 - 775.", "JAEGER, M. M., E. HAQUE, P. SULTANA, AND R. L. BRUGGERS. 2007. Daytime cover, diet and space-use of golden jackals (Canis aureus) in agroecosystems of Bangladesh. Mammalia 71: 1 - 10.", "ROTEM, G., H. BERGER, R. KING, P. B. (KUTIEL) [SIC], AND D. SALTZ. 2011. The effect of anthropogenic resources on the space-use patterns of golden jackals. Journal of Wildlife Management 78: 132 - 136.", "MUKHERJEE, S., S. P. GOYAL, A. J. T. J OHNSINGH, AND M. R. P. LEITE PITMAN. 2004. The importance of rodents in the diet of jungle cat (Felis chaus), caracal (Caracal caracal) and golden jackal (Canis aureus) in Sariska Tiger Reserve, Rajasthan, India. Journal of Zoology (London) 262: 405 - 411.", "AMROUN, N., P. GIRAUDOUX, AND P. DELLATRE. 2006. A comparative study of the diets of two sympatric carnivores - the golden jackal (Canis aureus) and the common genet (Genetta genetta) - in Kabylia, Algeria. Mammalia 70: 247 - 254.", "BORKOWSKI, J., A. ZALEWSKI, AND R. MANOR. 2011. Diet composition of golden jackals in Israel. Annales Zoologici Fennici 48: 106 - 118.", "KRUUK, H. 1972. The spotted hyena. University of Chicago Press, Chicago, Illinois.", "ROSEVEAR, D. R. 1974. The carnivores of West Africa. Trustees of the British Museum (Natural History), London, United Kingdom.", "ROELKE- PARKER, M. E., ET AL. 1996. A canine distemper virus epidemic in Serengeti lions (Panthera leo). Nature 379: 441 - 445.", "ALEXANDER, K. A., P. W. KAT, R. K. WAYNE, AND T. K. FULLER. 1994. Serologic survey of selected canine pathogens among free-ranging jackals in Kenya. Journal of Wildlife Diseases 30: 486 - 491.", "SHAMIR, M., ET AL. 2001. Antibodies to selected canine pathogens and infestation with intestinal helminths in golden jackals (Canis aureus) in Israel. Veterinary Journal 162: 66 - 72.", "DUBEY, J. P., ET AL. 2010. Toxoplasmosis in sand cats (Felis margarita) and other animals in the Breeding Centre for Endangered Arabian Wildlife in the United Arab Emirates and Al Wabra Wildlife Preservation, the State of Qatar. Veterinary Parasitology 172: 195 - 203.", "CIROVIC, D., I. PAVLOVIC, A. PENEZIC, Z. KULIsIC, AND S. SELAKOVIC. 2015. Levels of infection of intestinal helminth species in the golden jackal Canis aureus from Serbia. Journal of Helminthology 89: 28 - 33.", "BLAGA, R., C. GHERMAN, D. SEUCOM, V. COZMA, AND P. BOIREAU. 2008. First identification of Trichinella sp. in golden jackal (Canis aureus) in Romania. Journal of Wildlife Diseases 44: 457 - 459.", "BERDYEVA, A. T. 1998. Dracunculiasis eradication in Turkmenistan. Country Report by the Ministry of Health, Turkmenistan.", "L. M. ISAEV INSTITUTE OF MEDICAL PARASITOLOGY. 1998. Dracunculiasis eradication in Uzbekistan. Country Report to the Ministry of Health, Uzbekistan.", "LAPINI, L., P. MOLINARI, L. DORIGO, G. ARE, AND P. BERALDO. 2009. Reproduction of the golden jackal (Canis aureus moreoticus I. Geoffroy Saint Hilaire, 1835) in Julian Pre-Alps, with new data on its range-expansion in the High-Adriatic Hinterland. Bollettino del Museo Civico di Storia Naturale di Venezia 60: 169 - 186.", "LAHMAR, S., ET AL. 2009. Echinococcus in the wild carnivores and stray dogs of northern Tunisia: the results of a pilot survey. Annals of Tropical Medicine & Parasitology 103: 323 - 331.", "PATTON, W. S. 1910. Preliminary report on a new piroplasm (Piroplasma gibsoni sp. nov.) found in the blood of the hounds of the Madras Hunt and subsequently discovered in the blood of the jackal Canis aureus. Bulletin de la Societe de Pathologie Exotique 3: 274 - 280.", "LANSZKI, J., M. HELTAI, AND L. SZABo. 2006. Feeding habits and trophic niche overlap between sympatric golden jackal (Canis aureus) and red fox (Vulpes vulpes) in the Pannonian ecoregion (Hungary). Canadian Journal of Zoology 84: 1647 - 1656.", "KINGDON, J. 1977. East African mammals. An atlas of evolution in East Africa. Volume IIIA: carnivores. Academic Press, London, United Kingdom.", "SINGH, A. N. 1983. A study of diverse prey species of python (Python molurus) with special reference to its interaction with jackal (Canis aureus). Tigerpaper 10: 31 - 32.", "MACDONALD, D. W. 1979. The flexible social system of the golden jackal, Canis aureus. Behavioral Ecology and Sociobiology 5: 17 - 38.", "MADDOCK, L. 1979. The \" migration \" and grazing succession. Pp. 104 - 129 in Serengeti, dynamics of an ecosystem (A. R. E. Sinclair and M. Norton-Griffiths, eds.). University of Chicago Press, Chicago, Illinois.", "COHEN, T. M., R. KING, A. D OLEV, A. B OLDO, A. L ICHTER- PELED, AND G. K. BAR- GAL. 2013. Genetic characterization of populations of the golden jackal and the red fox in Israel. Conservation Genetics 14: 55 - 63.", "JERDON, T. C. 1984. A hand book of the mammals of India. Mittal Publishers, Delhi, India.", "GALOV, A., ET AL. 2015. First evidence of hybridization between golden jackal (Canis aureus) and domestic dog (Canis familiaris) as revealed by genetic markers. Royal Society Open Science 2: 150450.", "BARDELEBEN, C., R. L. MOORE, AND R. K. WAYNE. 2005. A molecular phylogeny of the Canidae based on six nuclear loci. Molecular Phylogenetics and Evolution 37: 815 - 831.", "PERINI, F. A., C. A. M. RUSSO, AND C. G. SCHRAGO. 2010. The evolution of South American endemic canids: a history of rapid diversification and morphological parallelism. Journal of Evolutionary Biology 23: 311 - 322.", "DURANT, S., M. E. CRAFT, R. HILBORN, S. BASHIR, J. HANDO, AND L. THOMAS. 2011. Long-term trends in carnivore abundance using distance sampling in Serengeti National Park, Tanzania. Journal of Applied Ecology 48: 1490 - 1500.", "GIANNATOS, G., Y. MARINOS, P. MARAGOU, AND G. CATSADORAKIS. 2005. The status of the golden jackal (Canis aureus) in Greece. Belgian Journal of Zoology 135: 145 - 149."]} |
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Text |
author |
Moehlman, Patricia D. Hayssen, Virginia |
author_facet |
Moehlman, Patricia D. Hayssen, Virginia |
author_sort |
Moehlman, Patricia D. |
title |
Canis aureus Linnaeus 1758 |
title_short |
Canis aureus Linnaeus 1758 |
title_full |
Canis aureus Linnaeus 1758 |
title_fullStr |
Canis aureus Linnaeus 1758 |
title_full_unstemmed |
Canis aureus Linnaeus 1758 |
title_sort |
canis aureus linnaeus 1758 |
publisher |
Zenodo |
publishDate |
2018 |
url |
https://dx.doi.org/10.5281/zenodo.4591937 https://zenodo.org/record/4591937 |
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Indian Fuller Morrison Ferguson Stripe Bombay Sinclair Hodgson Osborn Lubbock Margarita Tera Gora Seitz Brus Markova Pitman |
geographic_facet |
Indian Fuller Morrison Ferguson Stripe Bombay Sinclair Hodgson Osborn Lubbock Margarita Tera Gora Seitz Brus Markova Pitman |
genre |
Canis lupus gray wolf |
genre_facet |
Canis lupus gray wolf |
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ftdatacite:10.5281/zenodo.4591937 2023-05-15T15:51:34+02:00 Canis aureus Linnaeus 1758 Moehlman, Patricia D. Hayssen, Virginia 2018 https://dx.doi.org/10.5281/zenodo.4591937 https://zenodo.org/record/4591937 unknown Zenodo http://zenodo.org/record/4573460 http://publication.plazi.org/id/FFE8FF95AE11263519745504FF950F6E http://zoobank.org/9FC747F8-BAA4-4E2A-BC5B-53A20A0A03FA https://zenodo.org/communities/biosyslit https://dx.doi.org/10.1093/mspecies/sey002 http://zenodo.org/record/4573460 http://publication.plazi.org/id/FFE8FF95AE11263519745504FF950F6E https://dx.doi.org/10.5281/zenodo.4592352 https://dx.doi.org/10.5281/zenodo.4573462 https://dx.doi.org/10.5281/zenodo.4573464 http://zoobank.org/9FC747F8-BAA4-4E2A-BC5B-53A20A0A03FA https://dx.doi.org/10.5281/zenodo.4591936 https://zenodo.org/communities/biosyslit Open Access Creative Commons Zero v1.0 Universal https://creativecommons.org/publicdomain/zero/1.0/legalcode cc0-1.0 info:eu-repo/semantics/openAccess CC0 Biodiversity Taxonomy Animalia Chordata Mammalia Carnivora Canidae Canis Canis aureus article-journal ScholarlyArticle Text Taxonomic treatment 2018 ftdatacite https://doi.org/10.5281/zenodo.4591937 https://doi.org/10.1093/mspecies/sey002 https://doi.org/10.5281/zenodo.4592352 https://doi.org/10.5281/zenodo.4573462 https://doi.org/10.5281/zenodo.4573464 https://doi.org/10.5281/zenodo.4591936 2022-03-10T10:39:52Z Canis aureus Linnaeus, 1758 Golden Jackal Canis aureus Linnaeus, 1758:40. Type locality “oriente;” restricted to “Benná Mts., Laristan, S. Persia ” by Thomas (1911:135). Lupus aureus Linnaeus, 1758:40. No type locality. From a pre- 1758 name. Vulpes indiae orientalis Linnaeus, 1758:41. No type locality. From a pre-1758 name. [ Canis ] anthus F. Cuvier, 1820:plate. Type locality “ Senegal.” Canis variegatus Cretzschmar, 1826:31, plate 10. Type locality “Nubien. Oberes Egypten;” preoccupied. Canis syriacus Hemprich and Ehrenberg, 1830:sig z, plate 16. Type locality “e monte Lebano.” Canis lupaster Hemprich and Ehrenberg, 1833:sig ff. Type locality “Fayum, Egypt ” (not seen cited in Allen 1939:194). Canis sacer Hemprich and Ehrenberg, 1833:sig ff. Type locality “Fayum, Egypt ” (not seen cited in Allen 1939:194). Canis riparius Hemprich and Ehrenberg, 1833:sig ff. Type local- ity “Coast of Abyssinia, near Arkiko” (not seen cited in Allen 1939:195). Canis aureus indicus Hodgson, 1833:237. Type locality “Himalaya.” Canis aureus var. moreotica I. Geoffroy Saint-Hilaire, 1835:plate 1. Type locality “de Morée.” Thous anthus : Hamilton Smith, 1839:195. Name combination. Thous variegatus : Hamilton Smith, 1839:198. Name combination. Thous sengalensi Hamilton Smith, 1839:201, plate 13. Type locality “ Gambia and Senegal.” Sacalius aureus : Hamilton Smith, 1839:214. Name combination. Sacalius barbarus Hamilton Smith, 1839:218. Type locality “ Tunis;” preoccupied by Canis barbarus Shaw, 1800:311 synonym for Vulpes vulpes. Sacalius indicus : Hamilton Smith, 1839:219. Name combination. C [ anis ]. dalmatinus Wagner, 1841:383. No type locality; Dalmatia implied. Canis graecus Wagner, 1841:383. Type locality “Europaei.” Renaming of C. aureus moreotica I. Geoffroy Saint-Hilaire, 1835. C [ anis ]. aureus vulgaris Wagner, 1841:383. No type locality. Canis aureus algirensis Wagner, 1841:384. Type locality “d’Alger” from ‘chacal d’Alger’ I. Geoffroy Saint-Hilaire, 1835:22. C [ anis ]. aureus tripolitanus Wagner, 1841:384. No type locality; Tripoli, Tunis implied. Canis aureus typicus Kolenati, 1858:96. No type locality; Persia, Georgia, Turkey implied. Canis aureus typicus var caucasica Kolenati, 1858:96. No type locality; Caucasus implied. Canis aureus var syriaco Kolenati, 1858:96. Unauthorized emendation of syriacus Hemprich and Ehrenberg, 1830. Canis aureus var indica Kolenati, 1858:96. Unauthorized emendation of indicus Hodgson, 1833. Canis aureus var nubica Kolenati, 1858:96. Renaming of Canis variegatus Cretzschmar, 1826 but Kolenati attributes the name to Rüppell without giving a date. Canis aureus var algira Kolenati, 1858:96. Unauthorized emendation of algirensis Wagner, 1841. Canis aureus var senegalensis : Kolenati, 1858:96. Name combination. Canis aureus var barbarus : Kolenati, 1858:96. Name combination. Canis aureus balcanicus Brusina, 1892:317. Type locality “Nardu — nadomak Valpova — na obali Drave, Slavonija.” Canis hadramaticus Noack, 1896:356. Type locality “Hadramaut in Arabien.” A jackal-wolf composite. The jackal was chosen as the lectotype (Morrison-Scott 1939) Canis mengesi Noack, 1897:518. Type locality “Ostafrikanische.” Canis lupus minor Mojsisovico, 1897:202, 241, 244. Preoccupied by Canis spelaeus minor Wagner, 1829 according to Kretzoi, 1947. Canis cruesemanni Matschie, 1900:145. Type locality “ Siam.” Canis anthus soudanicus Thomas, 1903:295. Type locality “El Obeid,” Kordofan, Sudan. Canis somalicus Lorenz, 1906:306. Type locality “Ireso bei Agada.” Canis gallaënsis Lorenz, 1906:307. Type locality “Ginea (Arussi).” Canis doederleini Hilzheimer, 1906a:116. Type locality “Oberägypten.” Canis thooides Hilzheimer, 1906b:364. Type locality “Sennaar.” Canis lupaster grayi Hilzheimer, 1906b:367. Type locality “Nordafrika.” Canis studeri Hilzheimer, 1906b:368. Type locality “ Tunis.” C [ anis ]. mengesi lamperti Hilzheimer, 1906b:371. No type locality. Thos aureus bea Heller, 1914:5. Type locality “Loita Plains, British East Africa,” Kenya. Canis indicus kola Wroughton, 1916:651. Type locality “Palanpur,” India. Canis naria Wroughton, 1916:651. Type locality “Coorg,” west coast of India. Canis lanka Wroughton, 1916:652. Type locality “Mankeni,” Sri Lanka. Thos lupaster maroccanus Cabrera, 1921:263. Type locality “ Mogador (Marruecos).” Th [ os ]. l [ upaster ]. Algirensis : Cabrera, 1921:263. Name combination. Th [ os ]. a [ ureus ]. Riparius : Cabrera, 1921:264. Name combination. Th [ os ]. aureus nubianus Cabrera, 1921:264. Renaming of variegatus Cretzschmar, 1826. Canis aureus hungaricus Éhik, 1938:11. Type locality “Tyukod (Marshy-land Ecsed), Comitat Szatmár, Hungary;” Preoccupied according to Kretzoi, 1947. Canis aureus minor : Éhik, 1938:13. Name combination; preoccupied. Thos aureus algirensis : Allen, 1939:194. Name combination. Thos aureus anthus : Allen, 1939:194. Name combination. Thos aureus lupaster : Allen, 1939:194. Name combination. Thos aureus maroccanus : Allen, 1939:195. Name combination. Thos aureus soudanicus : Allen, 1939:195. Name combination. Thos gallaensis : Allen, 1939:195. Name combination. Thos lamperti : Allen, 1939:195. Name combination. [ Canis aureus ] moreoticus Kretzoi, 1947:287. Unjustified emendation of moreotica I. Geoffroy Saint-Hilaire, 1835. Thos aureus ecsedensis Kretzoi, 1947:287. Renaming of C. a. hungaricus Éhik, 1938. Canis lupaster doederleini : Saleh and Basuony, 2014:49. Name combination. Canis aureus qattarensis Saleh and Basuony, 2014:49. Type locality “northern region of the Egyptian Western Desert.” CONTENT AND CONTEXT. Order Carnivora, family Canidae, subfamily Caninae, genus Canis . As many as 13 subspecies of C . aureus are distinguished through Europe and Africa (Allen 1939; Ellerman and Morrison-Scott 1951; Coetzee 1977). However, these subspecies may be part of a 3-species complex (see “Nomenclatural Notes”). Unfortunately, much of the literature does not specify geographic location or subspecies. Here, C . aureus is used for the global range from Africa into Eurasia (Wozencraft 2005). Therefore, we list the historical 13 subspecies of C. aureus (Heller 1914; Allen 1939; Ellerman and Morrison- Scott 1951; Coetzee 1977; Wozencraft 2005). The newly named subspecies, qattarensis Saleh and Basuony, 2014, is tentatively treated as a synonym of syriacus . Subspecies designations for the orientalis Linnaeus, 1758 and studeri Hilzheimer, 1906b are not known. C. a. algirensis Wagner, 1841. See above; synonyms are algira Kolenati, 1858; barbarus (Hamilton Smith, 1839); grayi Hilzheimer, 1906b; tripolitanus Wagner, 1841. C. a. anthus F. Cuvier, 1820. See above; synonym is senegalensis (Hamilton Smith, 1839). C. a. aureus Linnaeus, 1758. See above; synonyms are balcanicus Brusina, 1892; caucasica Kolenati, 1858; hadramauticus Noack, 1896; maroccanus (Cabrera, 1921); orientalis (Linnaeus, 1758); typicus Kolenati, 1858; vulgaris Wagner, 1841. C. a. bea (Heller, 1914). See above. C. a. cruesemanni Matschie, 1900. See above. C. a. ecsedensis (Kretzoi, 1947). See above; synonyms are hungaricus Ehik, 1938; minor Mojsisovico, 1897 (preoccupied). C. a. indicus Hodgson, 1833. See above; synonyms are indica Kolenati, 1858; kola Wroughton, 1916. C. a. lupaster Hemprich and Ehrenberg, 1833. See above; synonym is sacer Hemprich and Ehrenberg, 1833. C. a. moreotica I. Geoffroy Saint-Hilaire, 1835. See above; synonyms are dalmatinus Wagner, 1841; graecus Wagner, 1841. C. a. naria Wroughton, 1916. See above; synonym is lanka Wroughton, 1916. C. a. riparius Hemprich and Ehrenberg, 1833. See above; synonyms are gallaënsis Lorenz, 1906; hagenbecki Noack, 1897; lamperti Hilzheimer, 1906b; mengesi Noack, 1897; somalicus Lorenz, 1906. C. a. soudanicus Thomas, 1903. See above, synonyms are doederleini Hilzheimer, 1906a; nubianus (Cabrera, 1921); nubica Kolenati, 1858; thooides Hilzheimer, 1906b; variegatus Cretzschmar, 1826 (preoccupied). C. a. syriacus Hemprich and Ehrenberg, 1830. See above, synonym are qattarensis Saleh and Basuony, 2014; syriaco Kolenati, 1858. NOMENCLATURAL NOTES. The above-named list may represent 3 taxa of canids: Canis anthus (African wolf), C. aureus , and C. lupus (gray wolf). Genetic analyses of C . aureus and C. lupus suggest that C . aureus from Africa and Eurasia are distinct clades and should be considered to be separate species, that is C. anthus (Africa) and C. aureus (Eurasia—Koepfli et al. 2015). As noted above, the literature does not distinguish these species and we include both as C. aureus . In addition, large C . aureus in Egypt ( lupaster ) may be a cryptic subspecies of C . anthus (Rueness et al. 2011; Gaubert et al. 2012). Canis aureus lupaster (Hemprich and Ehrenberg 1833) is classified as a golden jackal; however Ferguson (1981) suggested that the taxon C. aureus lupaster , which is present in arid areas of Egypt and Libya (Osborn and Helmy 1980), may actually represent a small C. lupus rather than a large jackal. Mitochondrial DNA analyses suggest that C. a. lupaster represents an ancient wolf lineage that may have colonized Africa prior to the northern hemisphere radiation (Rueness et al. 2011). Mitochondrial DNA analyses assigned individuals from Algeria, Mali, and Senegal to C . lupus lupaster (Gaubert et al. 2012) and identified C. lupus lupaster mtDNA haplotypes in C. aureus from Senegal questioning the genetic differentiation between the proposed C . anthus and C . aureus . We retain C. aureus lupaster as a form of the golden jackal not as a subspecies of C. lupus following Wozencraft (2005— Qumsiyeh 1996; Ferguson 2002). The specific name is Latin for golden. Common names include Asiatic jackal, common jackal, ibn awa or ibn awee (classic Arab), wa wie (spoken Arab), cakalli (Albanian), čagalj (Croatian), šakal obecný (Czech), sjakal (Danish, Swedish), jakhal (Dutch), šaakal (Estonian), sakaali (Finnish), Sjakalur (Faeroese), chacal doré (French), Goldschakal (German), tơɑκɑλ (Greek), aranysakál (Hungarian), sciacallo dorato (Italian), chagal, turg (Kurdish), zeltainais šakālis (Latvian), xakali (Maltese), gullsjakal (Nowegian), shoghal (Persian), szakal zlocisty (Polish), chacal-dourado (Portuguese), şakal (Rumanian), šakal obyčajný (Slovakian, Slovenian), chacal (Spanish), çakal (Turkish—Hatt 1959; P. D. Moehlman, in litt.). Indigenous names include Amharic: tera kebero (Ethiopia); Fulani: sundu; Hausa: dila; Hindi: Giddhad; Kanada: nuree; Kiswahili: bweha wa mbugani, bweha dhahabu (Tanzania); Marathi (India): kolha; Nepali (Nepal), Bengali, Gujarati and Kutchi (India): shiyal; Singhelese: nariya; Songhai: nzongo; Tamil (India): peria naree; Wolof: Tili (P. D. Moehlman, in litt.). DIAGNOSIS Canis aureus occurs sympatrically with the silver-backed jackal ( Canis mesomelas ) and the side-striped jackal ( Canis adustus ) in parts of East Africa. All have skull lengths of 141– 147 mm with minimal divergence in size (van Valkenburgh and Wayne 1994). C . mesomelas is distinguished by reddish flanks and limbs, and the skull may have a less developed parietal crest (Clutton-Brock et al. 1976), the angle of its ears which are more upright, and, usually, the prominent dark “silver” saddle composed of black and white hairs (Walton and Joly 2003). C . aureus may have a dark saddle seasonally, which is not prominent. C . adustus characteristically has shorter ears set more to the side of the skull, a pale side-stripe, and a white-tipped tail (Moehlman and Jhala 2013). C. lupus is much larger with a heavier body and variable pelage coloring. The skull is much larger with a welldeveloped interparietal crest, strong jaw, and large canine and carnassial teeth (Clutton-Brock et al. 1976). GENERAL CHARACTERS Canis aureus (Fig. 1) is a medium-sized canid, considered the most typical representative of the genus Canis (Clutton- Brock et al. 1976). Basic coat color is golden but varies from pale creamy yellow to a dark tawny hue on a seasonal basis. The pelage on the back is often a mixture of black, brown, and white hairs, such that they can appear to have a dark saddle similar to the C. mesomelas (Walton and Joly 2003; Moehlman and Jhala 2013). The long contour hairs have an extensive black tip and a wider penultimate white band (Pocock 1938). In winter, a copious under wool is present (Pocock 1938). C . aureus inhabiting rocky, mountainous terrain may have a grayer coat shade (Sheldon 1992). The belly and underparts are a lighter pale-ginger to cream. The tail is bushy with a tan to black tip. Females have 7–8 mammae (Poché et al. 1987), comprising 4 pairs (Sheldon 1992). Eye color varies from pale yellow to amber (Lewis et al. 1968). Means and parenthetical ranges of external measurements (mm) from 6 males and 3 females, respectively, in Gujarat, India (Y. V. Jhala, in litt.) were: length of head and body, 793 (760–840), 760 (740–800); length of tail, 220 (200–240), 205 (200–210); length of ear, 76 (68–90), 80 (75–85); body mass (kg) of 6 male and 4 females was 8.8 (7.6–9.8) and 7.3 (6.5– 7.8), respectively. In Tanzania, the external measurements (mm or kg) from 2 males in the Serengeti (Moehlman and Jhala 2013) were: length of head and body, 740, 785; length of tail, 270, 280; length of ear, 110, 110; mass, 6.3, 7.7. In Egypt, means and parenthetical ranges (mm) for 9 adult C. a. lupaster of unknown sex were: length of head and body, 872 (822–893); length of tail, 312 (290–347); length of hind foot, 200 (190–212); length of ear, 112 (104–121); and body mass (kg) of 4 adults was 13 (10–15—Osborn and Helmy 1980). In Lebanon, external measurements (mm) of 3 adult C. a. syriacus were: total length, 988.3 (950–1015); length of tail, 275 (265–290); length of head and body, 713.3 (680–750); length of hind foot, 156 (140–170); length of ear from notch, 74.7 (70–80— Lewis et al. 1968). In Bangladesh, mean measurements (mm, SD , n ) of females and males, respectively, were: total length, 1009.3 (33.67, 6), 1058.6 (20.30, 7); length of tail, 220.1 (17.16, 7), 245.0 (21.41, 6); length of hind paw, 58.0 (4.93, 7), 56.3 (4.83, 7); length of hind foot, 146.2 (24.66, 5), 162.6 (8.52, 7); length of ear, 69.4 (4.80, 5), 74.1 (5.49, 7); mass (kg), 8.5 (0.73, 7), 10.3 (0.95, 7— Poché et al. 1987). Mean adult mass (kg) of 5 females and 3 males, respectively, was 10.0 and 12.0 (Palmqvist et al. 2002). On average, females are 12% lighter than males: 5.8 kg versus 6.6 kg (Moehlman and Hofer 1997). Mean cranial measurements (mm, SD Fig. 2) on 10 mandibles and upper jaws of adult C . aureus in Iran were: length of skull, 169, 31.9; length of cranium, 101, 23.7; nasal length, 67, 16.6; cranial width, 72, 4.4; length of mandible, 112, 34.6; lateral alveolar root to mental foramen, 25, 1.7; mental foramen to caudal mandibular border, 87, 3.8; mandibular foramen to base of mandible, 12, 0.4; caudal border of mandible to below mandibular foramen, 11, 0.8; maximum mandibular height, 60, 5.8 (Mofared 2013). Mean measurements (mm, range) for 3 Lebanese adults were: greatest length of skull, 148.6 (135.2–157.4); condylobasal length, 142.9 (131.9–149.1); zygomatic breadth, 83.9 (73.3–90.2); breadth of braincase, 48.8 (46.0–50.3); interorbital constriction, 29.4 (25.5–32.7); maxillary toothrow, 62.9 (60.4– 65.0); mandibular toothrow, 72.2 (58.2–74.9); mandibular length, 114.1 (104.1–121.0— Lewis et al. 1968). Mean cranial measurements (mm; with range and n ) of Egyptian adult C. a. lupaster were: condyloincisive length, 185.2 (173.5–196.0, 13); zygomatic width, 101.4 (93.5–111.3, 14); rostral width, 33.8 (31.2–37.8, 14); postorbital width, 34.8 (31.8–38.9, 13); basioccipital width at tempoparietal suture, 54.4 (51.8–59.1, 13); nasal length, 72.0 (65.9–84.8, 14); width across upper molars, 58.2 (54.2–63.0, 14); distance from anterior most surface of canine to posterior of 2nd molar, 80.3 (75.6–86.8, 14); skull height, 66.8 (62.0–74.0, 9—Osborn and Helmy 1980). In Bulgaria, mean cranial and tooth measurements (mm) for 92 males and 56 females, respectively, were: total skull length, 166, 161; neurocranium length, 89, 86.5; greatest length of nasals, 59.7, 57.9; greatest neurocranium breath, 52.5, 52.6; total length of mandible, 121.9, 118.6; height of vertebral ramus, 48.6, 46.7; length of P1 to P4, 33.9, 33.4; length of M1 to M3, 31.8, 31.0 (Stoyanev 2012). Mean mandible length and depth at 2nd molar (mm) for 5 females and 3 males, respectively, was f: 102.8, 14.3; m: 108.9, 15.3 (Palmqvist 2002). Condylobasal length (mm) from 10 males was 154, whereas it was 140 from 7+ female skulls (Pocock 1938). DISTRIBUTION Canis aureus is widespread in North and East Africa, Europe, the Middle East, Central Asia, and Southeast Asia (Fig. 3). They range from sea level in Eritrea to 3,500 m in the Bale Mo : Published as part of Moehlman, Patricia D. & Hayssen, Virginia, 2018, Canis aureus (Carnivore: Canidae), pp. 14-25 in Mammalian Species 50 (957) on pages 14-22, DOI: 10.1093/mspecies/sey002, http://zenodo.org/record/4573460 : {"references": ["LINNAEUS, C. 1758. Systema naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Editio decima, reformata. Volume 1. Laurentii Salvii, Stockholm, Sweden.", "CUVIER, F. 1820. Le chacal du Senegal, femelle. Pp. 1 - 3 in Histoire naturelle des mammiferes, avec des figures dessinees d'apres les animaux vivants, tome duexieme (E. Geoffroy Saint-Hilaire and F. Cuvier, eds.). A. Belin, Paris, France. 1824 on title page but material on the jackal is June 1820.", "CRETZSCHMAR, P. J. 1826. Atlas zu der Reise im nordlichen Afrika von Eduard Ruppell. Erste Abtheilung Zoologie. Gedruckt und in Commission bei Heinr. Ludw. Bronner, Frankfurt, Germany.", "ALLEN, G. M. 1939. A checklist of African mammals. Bulletin of the Museum of Comparative Zoology at Harvard College 83: 1 - 763.", "HODGSON, B. H. 1833. Description of the wild dog of the Himalaya. Asiatic Research 18: 221 - 237.", "HAMILTON SMITH, C. 1839. The natural history of dogs: Canidae or genus Canis of authors; including also the genera Hyaena and Proteles. Pp. 77 - 267 in Jardine's naturalist library. Volume 18. Mammalia. Dogs. Volume 1 (W. Jardine, ed.). W. H. Lizars, Edinburgh, Scotland.", "SHAW, G. 1800. General zoology or systematic natural history. Volume 1, part 2. G. Kearsley, London, United Kingdom.", "KOLENATI, F. A. 1858. Reiseerinnerungen. Die Bereisung Hocharmeniens und Elisabethopols, der Schekinschen Provinz und des Kasbek im Central-Kaukasus. Verlagsbuchhandlung von Rudolf Kuntze, Dresden, Germany.", "BRUSINA, S. 1892. Cagalj balkanski (Canis aureus balcanicus Brus n. form.) iz Slavonije. Glasnik Hrvatskoga Naravoslovnoga Drustva Zagreb 7: 316 - 317.", "NOACK, T. 1896. I. Wissenschaftliche Mittheilungen. 1. Ein neuer Steinbock und ein neuer (?) [sic] Canide aus Arabien. Zoologischer Anzeiger 19: 353 - 356.", "NOACK, T. 1897. Ostafrikanische Schakale. Zoologischer Anzeiger 20: 517 - 521.", "MOJSISOVICO, A. 1897. Das Thierleben der Osterreichisch-Ungarischen Tiefebenen. Biologische und thiergeographische Skizzen und Bilder. A. Holder, Wien, Austria.", "KRETZOI, M. 1947. New names for mammals (Notes on nomenclature No. 3). Annales Historico-Naturales Musei Nationalis Hungarici 40: 285 - 287.", "MATSCHIE, P. 1900. Herr Matschie sprach uber den Schakal des Menam-Gebietes in Siam. Sitzungs-Berichte der Gesellschaft der Naturforschender Freunde zu Berlin 3: 144 - 145.", "LORENZ, L. 1906. Uber zwei neue Schakale aus Nordostafrika aus der Sammlung des Freiherrn C. von Erlanger. Anzeiger, Kaiserliche Akademie der Wissenschaften in Wien. Mathematisch- Naturwissenschaftliche Klasse 43: 306 - 310.", "HILZHEIMER, M. 1906 a. Papio mundamensis, Felis deliensis, Canis reissii und andre neue Saugetiere. Zoologischer Anzeiger 30: 109 - 118.", "HILZHEIMER, M. 1906 b. Die geographische Verbreitung der Afrikanischen Grauschakale. Zoologischer Beobachter 47: 363 - 373.", "HELLER, E. 1914. New subspecies of mammals from equatorial Africa. Smithsonian Miscellaneous Collections 63: 1 - 12.", "CABRERA, A. 1921. Algunos carnivoros africanos nuevos. Boletin de la Real Sociedad Espanola de Historia Natural, Madrid 21: 261 - 264.", "EHIK, J. 1938. Sakal vagy nadifarkas hazankbol [Jackal or reed-wolf from Hungary]. Annales Musei Nationalis Hungarici, Pars Zoological 31: 11 - 15.", "COETZEE, C. G. 1977. Order Carnivora. Part 8. Pp. 1 - 42 in The mammals of Africa: an identification manual (J. Meester and H. W Setzer, eds.). Smithsonian Institution Press, Washington, D. C.", "RUENESS, E. K., ET AL. 2011. The cryptic African wolf: Canis aureus lupaster is not a golden jackal and is not endemic to Egypt. PLoS One 6: e 16385.", "GAUBERT, P., ET AL. 2012. Reviving the African wolf Canis lupus lupaster in North and West Africa: a mitochondrial lineage ranging more than 6,000 km wide. PLoS One 7: e 42740.", "FERGUSON, W. W. 1981. The systematic position of Canis aureus lupaster (Carnivora: Canidae) and the occurrence of Canis lupus in North Africa, Egypt and Sinai. Mammalia 45: 459 - 465.", "QUMSIYEH, M. B. 1996. Mammals of the Holy Land. Texas Tech University Press, Lubbock.", "FERGUSON, W. W. 2002. The mammals of Israel. Gefen Publishing, Jeruslaem, Israel.", "CLUTTON- BROCK, J., G. B. CORBET, AND M. HILLS. 1976. A review of the family Canidae, with a classification by numerical methods. Bulletin of the British Museum (Natural History) Zoology 29: 1 - 199.", "POCOCK, R. I. 1938. The jackals of S. W. Asia and S. E. Europe. Proceedings of the Zoological Society of London 108: 37 - 40.", "SHELDON, J. W. 1992. Wild dogs: the natural history of the non-domestic Canidae. Academic Press, Chicago, Illinois.", "POCHE, R. M., S. J. EVANS, P. SULTANA, M. E. HAGUE, R. STERNER, AND M. A. SIDDIQUE. 1987. Notes on the golden jackal (Canis aureus) in Bangladesh. Mammalia 51: 259 - 270.", "LEWIS, R. E., J. H. LEWIS, AND S. I. ATALLAH. 1968. A review of Lebanese mammals. Carnivora, Pinnipedia, Hyracoidea and Artiodactyla. Journal of Zoology (London) 154: 517 - 531.", "PALMQVIST, P., M. MENDOZA, A. ARRIBAS, AND D. R. GRoCKE. 2002. Estimating the body mass of Pleistocene canids: discussion of some methodological problems and a new ' taxon free' approach. Lethaia 35: 358 - 360.", "MOFARED, A. L. 2013. Macro-anatomical investigation of the skull of golden jackal (Canis aureus) and its clinical application during regional anesthesia. Global Veterinaria 10: 547 - 550.", "STOYANEV, S. 2012. Craniometric differentiation of golden jackals (Canis aureus L., 1758) in Bulgaria. Pp. 39 - 47 in International symposium on hunting, modern aspects of sustainable management of game populations, Zemun-Belgrade, Serbia.", "PRATER, S. 1971. The book of Indian animals. Bombay Natural History Society, Bombay, India.", "ADMASU, E., S. J. THIRGOOD, A. BEKELE, AND K. M. LAURENSON. 2004. Spatial ecology of golden jackal in farmland in the Ethiopian highlands. African Journal of Ecology 42: 144 - 152.", "MALEZ, V. 1984. The zooarcheological data as the base of colonizing the Markova cave on the island of Hvar. Pp. 617 - 621 in 9 th Yugoslavian speleological congress proceedings, Yugoslavia (in Croatian).", "RUTKOWSKI, R., ET AL. 2015. A European concern? Genetic structure and expansion of golden jackals (Canis aureus) in Europe and the Caucasus. PLoS One 10: e 0141236.", "FABBRI, E., ET AL. 2014. Genetic structure and expansion of golden jackals (Canis aureus) in the north-western distribution range (Croatia and eastern Italian Alps). Conservation Genetics 15: 187 - 199.", "ARNOLD, J., A. HUMER, M. HELTAI, D. MURARIU, N. SPASSOV, AND K. HaCKLANDER. 2012. Current status and distribution of golden jackals Canis aureus in Europe. Mammal Review 42: 1 - 11.", "KRYsTUFEK, B., D. MURARIU, AND C. KURTONUR. 1997. Present distribution of the golden jackal Canis aureus in the Balkans and adjacent regions. Mammal Review 27: 109 - 114.", "HOOIJER, D. A. 1961. The fossil vertebrates of Ksar'Akil, a Paleolithic rockshelter in Lebanon. Zoologische Verhandelingen 49: 1 - 68.", "SAVAGE, R. J. 1977. Evolution in carnivorous mammals. Paleontology 20: 237 - 271.", "KURTEN, B. 1974. A history of coyote-like dogs (Canidae, Mammalia). Acta Zoologica Fennica 140: 1 - 38.", "SPASSOV, N. 1989. The position of jackals in the Canis genus and lifehistory of the golden jackal (Canis aureus L.) in Bulgaria and the Balkans. Historia Naturalis Bulgarica, Sofia 1: 44 - 56.", "EWER, R. F. 1956. The fossil carnivores of the Transvaal caves: Canidae. Proceedings of the Zoological Society of London 126: 97 - 120.", "DIXSON, A. F. 1995. Baculum length and copulatory behaviour in carnivores and pinnipeds (Grand Order Ferae). Journal of Zoology (London) 235: 67 - 76.", "FULLER, T. K., A. R. BIKNEVICIUS, P. W. KAT, B. VAN VALKENBURGH, AND R. K. WAYNE. 1989. The ecology of three sympatric jackal species in the Rift Valley of Kenya. African Journal of Ecology 27: 313 - 323.", "SEITZ, V. A. 1959. Beobachtungen an Handaufgezogenen Goldschakalen (Canis aureus aligirensis). Zeitschrift fur Tierpsychologie 16: 747 - 771.", "MOEHLMAN, P. D. 1983. Socioecology of silverbacked and golden jackals (Canis mesomelas and Canis aureus). Pp. 423 - 453 in Advances in the study of mammalian behavior (J. F. Eisenberg and D. G. Kleiman, eds.). American Society of Mammalogists Special Publications, Lawrence, Kansas.", "MOEHLMAN, P. D. 1986. Ecology of cooperation in canids. Pp. 64 - 86 in Ecological aspects of social evolution (D. I. Rubenstein and R. W. Wrangham, eds.). Princeton University Press, Princeton, New Jersey.", "RAICHEV, E. 2011. Effect of shooting on the structure of population of golden jackal (Canis aureus L.) in Sarnena Sredna Gora Mountain. Agricultural Science and Technology 3: 276 - 280.", "FLOWER, S. S. 1932. Notes on the recent mammals of Egypt, with a list of the species recorded from that kingdom. Proceedings of the Zoological Society of London 1932: 369 - 450.", "Ginsberg, J. R., AND D. W. Macdonald (eds.). 1990. Foxes, wolves, jackals, and dogs. An action plan for the conservation of canids. International Union for the Conservation of Nature, Gland, Switzerland.", "MOEHLMAN, P. D. 1989. Intraspecific variation in canid social systems. Pp. 143 - 163 in Carnivore behavior, ecology, and evolution (J. L. Gittleman, ed.). Chapman & Hall, London, United Kingdom.", "KROFEL, M. 2008. Survey of golden jackals (Canis aureus L.) in northern Dalmatia, Croatia: preliminary results. Natura Croatia 17: 259 - 264.", "KROFEL, M. 2007. Golden jackals (Canis aureus L.) on the Peljesac Peninsula (southern Dalmatia, Croatia). Natura Croatica 16: 201 - 204.", "KHIDAS, K. 1990. Contribution a la connaissance du chacal dore. Facteurs modulant l'organisation sociale et territorial de la sous-espece algerienne (Canis aureus algirensis Wagner, 1841). Mammalia 54: 361 - 375.", "JAEGER, M. M., R. K. PANDIT, AND E. HAQUE. 1996. Season differences in territorial behavior by golden jackals in Bangladesh: howling versus confrontation. Journal of Mammalogy 77: 768 - 775.", "JAEGER, M. M., E. HAQUE, P. SULTANA, AND R. L. BRUGGERS. 2007. Daytime cover, diet and space-use of golden jackals (Canis aureus) in agroecosystems of Bangladesh. Mammalia 71: 1 - 10.", "ROTEM, G., H. BERGER, R. KING, P. B. (KUTIEL) [SIC], AND D. SALTZ. 2011. The effect of anthropogenic resources on the space-use patterns of golden jackals. Journal of Wildlife Management 78: 132 - 136.", "MUKHERJEE, S., S. P. GOYAL, A. J. T. J OHNSINGH, AND M. R. P. LEITE PITMAN. 2004. The importance of rodents in the diet of jungle cat (Felis chaus), caracal (Caracal caracal) and golden jackal (Canis aureus) in Sariska Tiger Reserve, Rajasthan, India. Journal of Zoology (London) 262: 405 - 411.", "AMROUN, N., P. GIRAUDOUX, AND P. DELLATRE. 2006. A comparative study of the diets of two sympatric carnivores - the golden jackal (Canis aureus) and the common genet (Genetta genetta) - in Kabylia, Algeria. Mammalia 70: 247 - 254.", "BORKOWSKI, J., A. ZALEWSKI, AND R. MANOR. 2011. Diet composition of golden jackals in Israel. Annales Zoologici Fennici 48: 106 - 118.", "KRUUK, H. 1972. The spotted hyena. University of Chicago Press, Chicago, Illinois.", "ROSEVEAR, D. R. 1974. The carnivores of West Africa. Trustees of the British Museum (Natural History), London, United Kingdom.", "ROELKE- PARKER, M. E., ET AL. 1996. A canine distemper virus epidemic in Serengeti lions (Panthera leo). Nature 379: 441 - 445.", "ALEXANDER, K. A., P. W. KAT, R. K. WAYNE, AND T. K. FULLER. 1994. Serologic survey of selected canine pathogens among free-ranging jackals in Kenya. Journal of Wildlife Diseases 30: 486 - 491.", "SHAMIR, M., ET AL. 2001. Antibodies to selected canine pathogens and infestation with intestinal helminths in golden jackals (Canis aureus) in Israel. Veterinary Journal 162: 66 - 72.", "DUBEY, J. P., ET AL. 2010. Toxoplasmosis in sand cats (Felis margarita) and other animals in the Breeding Centre for Endangered Arabian Wildlife in the United Arab Emirates and Al Wabra Wildlife Preservation, the State of Qatar. Veterinary Parasitology 172: 195 - 203.", "CIROVIC, D., I. PAVLOVIC, A. PENEZIC, Z. KULIsIC, AND S. SELAKOVIC. 2015. Levels of infection of intestinal helminth species in the golden jackal Canis aureus from Serbia. Journal of Helminthology 89: 28 - 33.", "BLAGA, R., C. GHERMAN, D. SEUCOM, V. COZMA, AND P. BOIREAU. 2008. First identification of Trichinella sp. in golden jackal (Canis aureus) in Romania. Journal of Wildlife Diseases 44: 457 - 459.", "BERDYEVA, A. T. 1998. Dracunculiasis eradication in Turkmenistan. Country Report by the Ministry of Health, Turkmenistan.", "L. M. ISAEV INSTITUTE OF MEDICAL PARASITOLOGY. 1998. Dracunculiasis eradication in Uzbekistan. Country Report to the Ministry of Health, Uzbekistan.", "LAPINI, L., P. MOLINARI, L. DORIGO, G. ARE, AND P. BERALDO. 2009. Reproduction of the golden jackal (Canis aureus moreoticus I. Geoffroy Saint Hilaire, 1835) in Julian Pre-Alps, with new data on its range-expansion in the High-Adriatic Hinterland. Bollettino del Museo Civico di Storia Naturale di Venezia 60: 169 - 186.", "LAHMAR, S., ET AL. 2009. Echinococcus in the wild carnivores and stray dogs of northern Tunisia: the results of a pilot survey. Annals of Tropical Medicine & Parasitology 103: 323 - 331.", "PATTON, W. S. 1910. Preliminary report on a new piroplasm (Piroplasma gibsoni sp. nov.) found in the blood of the hounds of the Madras Hunt and subsequently discovered in the blood of the jackal Canis aureus. Bulletin de la Societe de Pathologie Exotique 3: 274 - 280.", "LANSZKI, J., M. HELTAI, AND L. SZABo. 2006. Feeding habits and trophic niche overlap between sympatric golden jackal (Canis aureus) and red fox (Vulpes vulpes) in the Pannonian ecoregion (Hungary). Canadian Journal of Zoology 84: 1647 - 1656.", "KINGDON, J. 1977. East African mammals. An atlas of evolution in East Africa. Volume IIIA: carnivores. Academic Press, London, United Kingdom.", "SINGH, A. N. 1983. A study of diverse prey species of python (Python molurus) with special reference to its interaction with jackal (Canis aureus). Tigerpaper 10: 31 - 32.", "MACDONALD, D. W. 1979. The flexible social system of the golden jackal, Canis aureus. Behavioral Ecology and Sociobiology 5: 17 - 38.", "MADDOCK, L. 1979. The \" migration \" and grazing succession. Pp. 104 - 129 in Serengeti, dynamics of an ecosystem (A. R. E. Sinclair and M. Norton-Griffiths, eds.). University of Chicago Press, Chicago, Illinois.", "COHEN, T. M., R. KING, A. D OLEV, A. B OLDO, A. L ICHTER- PELED, AND G. K. BAR- GAL. 2013. Genetic characterization of populations of the golden jackal and the red fox in Israel. Conservation Genetics 14: 55 - 63.", "JERDON, T. C. 1984. A hand book of the mammals of India. Mittal Publishers, Delhi, India.", "GALOV, A., ET AL. 2015. First evidence of hybridization between golden jackal (Canis aureus) and domestic dog (Canis familiaris) as revealed by genetic markers. Royal Society Open Science 2: 150450.", "BARDELEBEN, C., R. L. MOORE, AND R. K. WAYNE. 2005. A molecular phylogeny of the Canidae based on six nuclear loci. Molecular Phylogenetics and Evolution 37: 815 - 831.", "PERINI, F. A., C. A. M. RUSSO, AND C. G. SCHRAGO. 2010. The evolution of South American endemic canids: a history of rapid diversification and morphological parallelism. Journal of Evolutionary Biology 23: 311 - 322.", "DURANT, S., M. E. CRAFT, R. HILBORN, S. BASHIR, J. HANDO, AND L. THOMAS. 2011. Long-term trends in carnivore abundance using distance sampling in Serengeti National Park, Tanzania. Journal of Applied Ecology 48: 1490 - 1500.", "GIANNATOS, G., Y. MARINOS, P. MARAGOU, AND G. CATSADORAKIS. 2005. The status of the golden jackal (Canis aureus) in Greece. 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