Homolophus arcticus Banks 1893
Homolophus arcticus Banks, 1893 Figures 10–11 Homolophus arcticus Banks, 1893: 208; Staręga 1978: 209; Cokendolpher 1987: 91; Tsurusaki 1987: 97–102, figs. 1–5; Li & Song 1993: 239; Crawford (1996 web page); Zhu et al. 1999: 7–8, fig. 1; Tchemeris 2000: 35–36, figs. 12–16; Tsurusaki &a...
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2021
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Online Access: | https://dx.doi.org/10.5281/zenodo.4450791 https://zenodo.org/record/4450791 |
Summary: | Homolophus arcticus Banks, 1893 Figures 10–11 Homolophus arcticus Banks, 1893: 208; Staręga 1978: 209; Cokendolpher 1987: 91; Tsurusaki 1987: 97–102, figs. 1–5; Li & Song 1993: 239; Crawford (1996 web page); Zhu et al. 1999: 7–8, fig. 1; Tchemeris 2000: 35–36, figs. 12–16; Tsurusaki & Song 2000: 162, fig. 5; Staręga 2003: 96 Opilio sachaliensis Suzuki, 1941: 245–247, figs. 8–13; Suzuki 1956: 97–100, figs. 1–7 [junior subjective synonym of Phalangium nordeskiöldi var. albofasciatum Kulczyński, 1901 by Staręga (1964); removed to synonymy of Egaenus zichyi Kulczyński 1901 by Staręga (1978); synonymies rejected and considered as junior subjective synonym of Homolophus arcticus Banks, 1893 by Tsurusaki (1987)]. Opilio sunuitensis Nakatsudi, 1943: 108–109, fig. 2 Opilio sachalinus Roewer, 1956: 277, 295, pl. 41, figs. 153–155 [junior subjective synonym of Phalangium nordeskiöldi var. albofasciatum Kulczyński, 1901 by Staręga (1964); removed to synonymy of Egaenus zichyi Kulczyński 1901 by Staręga (1978); synonymies rejected and considered as junior subjective synonym of Homolophus arcticus Banks, 1893 by Cokendolpher (1987)]. Opilio kurilus Roewer, 1956: 279, 280–289, pl. 40, figs. 131–133 [junior subjective synonym of Phalangium nordeskiöldi var. albofasciatum Kulczyński, 1901 by Staręga (1964); removed to synonymy of Egaenus zichyi Kulczyński 1901 by Staręga (1978); synonymies rejected and considered as junior subjective synonym of Homolophus arcticus Banks, 1893 by Tsurusaki (1987)]. Opilio nipponensis Roewer, 1956: 279, 292–293, pl. 40, figs. 139–141 [junior subjective synonym of Homolophus arcticus Banks, 1893 by Staręga (2003)] Euphalangium albofasciatum — Staręga 1964: 399 Egaenus zichyi — Staręga 1978: 222 Diagnosis . The species differs from all congeners, including the most closely related, H. chemerisi (Staręga & Snegovaya, 2008) and H. tibetanus (Roewer, 1911), by the following: cephalothorax with group of fairly large darklycolored-tipped denticles in center on anterior edge; short legs, I and III pairs thickened; penis glans thickened, with somewhat squared-off tip (lateral view) and oval (dorsal view); shape of truncus (wide at base gradually getting thinner to about distal third which stays slender to end). Type Locality . Russia, Kamchatka Oblast, Komandorskiye Ostrova- “Commander Island, Siberia”, ca. 55°12’0”N, 165°59’24”E. Distribution . China: Gansu, Xinjiang, Sichuan. Russia (Kamchatka Oblast and Sakhalin Oblast), Japan (Hokkaido), North Korea, and here newly recorded from Mongolia (Fig. 11). Specimens Examined . RUSSIA , Khabarovsk Territory, Bureinsky Reserve, birch-wood, 12– 17.08.2002, leg. L. Triliskauskas (1 male, 1 female, RCNS). MONGOLIA , 1899, leg. A. Soldatov, field # N151-1899 (1 male, 1 female, ZIN); Tov Aimak, 40 km ESE of Ulanbator, 47°48.88’N, 107°26.22’E, 1500 m a.s.l., 28– 30.07.2003, leg. O. Gorbunov (1 male, 4 females, 12 juv., RCNS). JAPAN, Wakasakanai, Toyotomi-chô, Teshio-gun, Hokkaido, 06.08.1985, leg. N. Tsurusaki (specimen already recorded by Tsurusaki, 1987) (1 male, AMNH). Description . Medium-sized harvestmen; male body rectangular, with rounded corners; medium sclerotization of body cuticle, less slender, 1.8 times longer than wide; length 6.5 mm, width 3.5 mm; with small hardly noticeable denticles along borders of tergites; cephalothorax darker and dark spots on abdomen arranged transversely along centers of tergites. Entire body length with central incomplete line of light color; Ocularium low, round to slightly wider than long, small, 1.9 times its length from anterior margin of cephalothorax, with 4 denticles on each side. Anterior edge of cephalothorax with group of fairly large darkly colored-tipped denticles; many smaller denticles in front of and lateral sides of area around ocularium. Legs short, angular in cross-section, I and III pairs thickened, especially Fe; Fe all legs with longitudinal rows of denticles. Leg lengths (mm): I 2.8 + 1.5 + 2.7 + 2.5 + 5.1 = 14.6, II 4.2 + 1.5 + 4.3 + 3.7 + 11.5 = 25.2, III 2.2 + 1.3 + 2.8 + 3.3 + 6.3 = 15.9, IV 4.2 + 1.4 + 3.2 + 4.8 + 9.0 = 22.6. Pedipalp segment lengths (mm): 1.5 + 0.8 + 1.0 + 1.5 = 4.8. Pedipalps thickened (sexually dimorphic), Fe ventrally and dorsally with denticles, Pa dorsally and laterally, Ti ventrally densely covered with denticles, dorsally and laterally are not densely covered, Ta ventrally covered by strip of sensory microdenticles on basal half. Chelicerae enlarged (sexually dimorphic) with some tabby marks, basal segment dorsally with several denticles, distal segment dorsally with few microdenticles. Cheliceral basal segment ventrally with blunt rounded bulge, length 2.0 mm, distal segment length 2.5 mm. Penis short, dorsoventrally flattened, bowed, recurved in lateral view, length (mm): truncus 2.9, glans 0.5, stylus 0.15; wide at base, gradually thinning on basal half of truncus, distal end uniform width extending to glans, glans oval in shape. Female differs from male in larger size, presence of densely placed denticles (few sharing same enlarged base) on anterior part of cephalothorax, in front of ocularium. Body rectangular, with rounded corners; less slender, 2.21 times longer than wide; body length 6.2 mm, width 2.8 mm. Leg lengths (mm): I 2.3 + 1.0 + 2.0 + 2.3 + 5.0 = 12.6, II 4.0 + 1.2 + 3.3 + 3.0 + 8.5 = 20.0, III 2.3 + 1.0 + 2.0 + 2.8 + 5.3 = 13.4, IV 3.5 + 1.3 + 2.7 + 4.3 + 7.6 = 19.4. Pedipalp segment lengths (mm): 1.0 + 0.5 + 0.8 + 1.3 = 3.6. Cheliceral basal segment length 1.4 mm, distal segment length 1.5 mm. Comments . The male studied from Japan differs slightly from illustrations herein by having the distal end of glans more truncated. Truncus of penis of Japanese specimen is very dorsoventrally flattened near middle and lower 1/2 of truncus and then more rounded in distal 1/4 end (see Tsurusaki 1987: fig. 4B). : Published as part of Snegovaya, Nataly Yu. & Cokendolpher, James C., 2021, Further studies on harvestman genus Homolophus (Opiliones: Phalangiidae), with descriptions of two new species, pp. 301-353 in Zootaxa 4908 (3) on pages 319-321, DOI: 10.11646/zootaxa.4908.3.1, http://zenodo.org/record/4441762 : {"references": ["Banks, N. (1893) The Phalanginae of the United States. The Canadian Entomologists, 25 (8), 205 - 211. https: // doi. org / 10.4039 / Ent 25205 - 8", "Starega, W. (1978) Katalog der Weberknechte (Opiliones) der Sowjet-Union. Fragmenta Faunistica, 23 (10), 197 - 241. Available from: https: // rcin. org. pl / dlibra / publication / 55993 (accessed 25 October 2020) https: // doi. org / 10.3161 / 00159301 FF 1978.23.10.197", "Cokendolpher, J. C. (1987) On the identity of the genus Homolophus: A senior synonym of Euphalangium (Opiliones: Phalangiidae). Acta Arachnologica, 35, 89 - 96. https: // doi. org / 10.2476 / asjaa. 35.89", "Tsurusaki, N. (1987) Two species of Homolophus newly found from Hokkaido, Japan (Arachnida: Opiliones: Phalangiidae). Acta Arachnologica, 35 (2), 97 - 107. Available from: https: // www. jstage. jst. go. jp / article / asjaa 1936 / 35 / 2 / 35 _ 2 _ 97 / _ article (accessed 25 October 2020) https: // doi. org / 10.2476 / asjaa. 35.97", "Li, S. - Q. & Song, D. - X. (1993) A revision of Chinese harvestmen, with a note on the geographical distribution (Arachnida: Opiliones). Scientific Treatise on Systematic and Evolutionary Zoology, 2 (4), 237 - 245. [in Chinese]", "Crawford, R. L. (1996) International Kuril Islands Project Expedition No. 3, July 25 - September 2, 1996. Field report on spider sampling: preliminary reconnaissance of the spider fauna of the northern Kuril Islands. Last viewed 12 March 2019. Available from: https: // www. burkemuseum. org / static / okhotskia / ikip / Results / reports / spider / rlcrpt 96. htm (accessed 25 October 2020)", "Zhu, M., Song, D. & Kim, J. - P. (1999) Notes on three species of the genus Homolophus (Opiliones: Phalangiidae) from China. Korean Arachnology, 15 (1), 7 - 12.", "Tchemeris, A. N. (2000) Contribution to the knowledge of the harvestman fauna in the Russian Far East and Eastern Siberia (Arachnida: Opiliones). Arthropoda Selecta, Moskva, 9 (1), 31 - 49. [figs. 17 - 20 not printed]", "Starega, W. (2003) On the identity and synonymies of some Asiatic Opilioninae (Opiliones: Phalangiidae). Acta Arachnologica, 52 (2), 91 - 102. https: // doi. org / 10.2476 / asjaa. 52.91", "Suzuki, S. (1941) Opiliones from southern Saghalien and Kaiba Island. Journal of Science of the Hiroshima University, Series B, Division 1 (Zoology), 9 (2), 239 - 248, pl. I.", "Suzuki, S. (1956) Further note on Opilio sachaliensis SUZUKI (Phalangida) from Southern Sakhalin and Southern Kuril Islands. Journal of Science of the Hiroshima University, Series B, Division 1 (Zoology), 16, 97 - 100.", "Kulczynski, W. (1901) Arachnoidea. In: Horvath, G. (Ed.), Dritte Asiatische Forschungsreise des Grafen Eugen Zichy. Vol. 2. Zoologische Ergebnisse. V. Hornyainszky, Budapest-Leipzig, pp. 311 - 369, pls. 12 - 13.", "Starega, W. (1964) Materialien zur Kenntnis der ostasiatischen Weberknechte (Opiliones). I-IV. Annales Zoologici [Polska Akademia Nauk], Warsawa, 22 (17), 387 - 410.", "Nakatsudi, K. (1943) [On some harvesters from Manchuria and Inner Mongolia]. Dobutsugaku Zasshi [Zoological Magazine], Tokyo, 55 (3), 106 - 113, figs. 1 - 15. [in Japanese with English summary]", "Roewer, C. F. (1956) Uber Phalangiinae (Phalangiidae, Opiliones Palpatores). (Weitere Weberknechte XIX). Senckenbergiana biologica, 37, 247 - 318.", "Roewer, C. F. (1911) Ubersicht der Genera der Subfamilie der Phalangiini der Opiliones Palpatores nebst Beschreibung einiger neuer Gattungen und Arten. Archiv fur Naturgeschichte, Berlin, Abt. A, Original-Arbeiten, 77 (Supplementheft 2), 1 - 106, pls. 1 - 3."]} |
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