Dryinus saussurei Ceballos 1936

• Main Authors: Olmi, Massimo, Copeland, Robert S., Noort, Simon Van
• Format: Text
• Language: unknown
• Published: Zenodo 2019
• Subjects: Biodiversity; Taxonomy; Animalia; Dryinus; Dryinus saussurei; Cape Walker; Izquierdo; Walker Bay
• Online Access: https://dx.doi.org/10.5281/zenodo.3511145; https://zenodo.org/record/3511145

31. Dryinus saussurei (Ceballos, 1936) (Figs 140 B–D) Lestodryinus saussurei Ceballos 1936: 48; Rey del Castillo & Izquierdo 1989: 254. Dryinus saussurei (Ceballos): Benoit 1954: 412; Olmi 1984: 770; 1994c: 10; 2006: 43; Azevedo et al . 2010: 891; Olmi et al . 2016: 76. Description. ♀. Fully winged; body length 4.0– 5.9 mm. Head black, except mandible and clypeus yellow; rarely clypeus black or brown; occasionally anterior region of frons testaceous; occasionally malar space with testaceous spot (in specimen from Botswana, Serowe); antenna testaceous; occasionally antennomeres 7–8, or 3–10, or 3–8 brown; propleuron black; pronotum black, with lateral regions and posterior collar reddish; rest of mesosoma black; metasoma black-brown; legs testaceous, except coxae and clubs of femora partly brown. Antenna clavate; antennomeres in following proportions: 10:5:21:10:9:8:6:6:4.5:8. Head (Fig. 140B) shiny or dull, completely or partly granulate, occasionally partly unsculptured; occasionally head partly granulate and partly sculptured by a few irregular keels; frontal line complete; occipital carina complete; lateral ocelli usually touching occipital carina, occasionally very close, but not touching occipital carina; OL = POL. In specimen from Madagascar, 17°45'S47°59'E, POL = 4; OL = 2. Pronotum dull, partly or completely granulate, with numerous slight striae around disc; occasionally pronotum shiny, almost completely unsculptured; posterior collar of pronotum short, about 0.5 × as long as distance from tegula to pronotal tubercle; pronotal tubercle not reaching tegula. Mesoscutum dull, slightly or strongly granulate, with short keels or areolae near posterior margin; occasionally median region of mesoscutum shiny, unsculptured (in specimen from Cameroon, Nkoemvon, mesoscutum completely granulate, with no areolae or keels; in specimen from South Africa posterior half of mesoscutum completely reticulate rugose; in specimen from Botswana, Serowe, mesoscutum almost completely reticulate rugose; in specimen from Cameroon, Messabengondo, mesoscutum completely granulate and rugose). Notauli incomplete, approximately reaching 0.65–0.90 × length of mesoscutum (rarely 0.5 × length of mesoscutum; rarely notauli complete and posteriorly separated). Mesoscutellum dull, granulate, or shiny and unsculptured. Metanotum dull, rugose. Metapectal-propodeal disc reticulate rugose; propodeal declivity reticulate rugose, with two longitudinal keels. Forewing with three brown transverse bands (two proximal bands almost joint, occasionally completely joint); distal part of 2r-rs&Rs vein about twice as long as proximal part (18:8). Hind wing hyaline, without dark spots; occasionally (in very large specimens) with one dark spot. Protarsomere 1 approximately as long as 4; in specimens from Madagascar, 19°07.56'S44°48.53'E and 18°01.35'S44°03.02'E, protarsomere 1 longer than 4; protarsomere 3 produced into hook. Enlarged claw (Fig. 140D) with one big subdistal tooth and one row of 9–13 lamellae (in specimen from Madagascar, 18°01.35'S44°03.02'E, subapical tooth distinctly divided into two teeth (Fig. 140C), so that apparently this specimen has to be included in group 3 of Dryinus ). Protarsomere 5 (Fig. 140D) with two rows of 16–39 lamellae; apex with 11–16 lamellae. Tibial spurs 1/1/2. ♂. Fully winged; body length 2.6–2.8 mm. Head black, except mandible testaceous; antenna brown, except scape and pedicel testaceous; mesosoma black; metasoma brown; legs testaceous, except coxae, club of metafemur and metatibia brown. Antenna filiform; antennomeres in following proportions: 8:7:17:12:13:13:11:10:10:11. Head dull, granulate; frons flat in front of anterior ocellus and humped between antennal toruli; frontal line incomplete; occipital carina incomplete; POL = 9; OL = 3; OOL = 3; lateral ocelli touching occipital carina; temple absent. Mesoscutum dull, granulate. Notauli incomplete, reaching approximately 0.65 × length of mesoscutum. Mesoscutellum almost completely dull, granulate, with narrow posterior region smooth and shiny. Metanotum with anterior half rugose and posterior half shiny and smooth. Metapectal-propodeal disc reticulate rugose, without transverse posterior keel; propodeal declivity reticulate rugose, without longitudinal keels. Forewing hyaline, without dark transverse bands; distal part of 2r-rs&Rs vein slightly longer than proximal part (14:13); 2R1 cell open. Paramere without dorsal process. Tibial spurs 1/1/2. Material examined . Type : ♀holotype: MADAGASCAR: Toamasina, Rogez (= Andekaleka), I.1931, A. Sey- rig leg. (MNMS). Other material : BOTSWANA: Central District, Serowe, XI.1985, MT, P. Forchhammer leg., 1♀ (NHMUK). CAMEROON: South Region, near Ebolowa, Nkoemvon, 30.XI.1979 – 5.I.1980, MT in forest, D. Jackson leg., NHMUK (E) 2007–19, 1♀ (NHMUK); same locality label, 23.IX–25.X.1980, 1♀ (NHMUK); Messaben- gondo, 1–15.III.2003, M. Tussac leg., 1♀ (HTS). ETHIOPIA: Shoa Prov., 95 km W Addis Ababa, Ambo, near Plant Protection Research Institute, 2400–2500 m, 19.VI.1990, A. Emel'yanov leg., 1♀ (ZISP). MADAGASCAR: Anta- nanarivo, La Mandraka, 1♀ (MRAC); same locality label, II.1953, Pierre Andria Robinson leg., 1♀ (MNHN); An- tananarivo, Analamanga Region, Ambatolaona, VI.1957, Pierre Andria Robinson leg., 1♀ (MNHN); Antananarivo, Andebadeba, 4 km from Arivanimamo, 19°00'29''S47°08'55''E, 1600 m, 7–14.X.2007, Uapaca forest, MT, M. Irwin, R. Harin'Hala leg., MA-46-02, 1♀ (CAS); Antananarivo, Ankokoy Forest, 3 km E of Ibity, 20°04.05'S46°59.97'E, 1700 m, 12–22.XII.2008, MT, in Uapaca forest, M. Irwin, R. Harin'Hala leg., MG-56-05, 1♀ (CAS); Antsiranana, Nosy Bé Island, 1♀ (HNHM); Antsiranana, Manongativo Reserve, 19.8 km 229° SW Antanambao, 13°58'S48°26'E, 400 m, 8–13.XI.1998, B.L. Fisher leg., 1♀ (DEUCD); Antsiranana, Marojejy National Park, 5 km W Manantenina village, Camp Mantella, 14°26.29'S49°46.44'E, 490 m, 28.IV–7.V.2005, MT, low altitude rainforest, M. Irwin & R. Harin'Hala leg., MA-31-20, 1♀ (CAS); Antsiranana, Marojejy National Park, Manantenina River, 27.6 km 35° NE Andapa, 9.6 km NNW Manantenina village, 14°26.06'S49°45.36'E, 775 m, 15–18.XI.2003, YPT, rainforest, BLF8873, B.L. Fisher et al . leg., 1♀ (CAS); Antsiranana, Marojejy National Park, Manantenina River, 28.0 km 38° NE Andapa, 8.2 km NNW Manantenina village, 14°26.12'S49°46.30'E, 450 m, 12–25.XI.2003, YPT, rainforest, BLF8725, B.L. Fisher et al . leg., 2♀♀ (CAS, MOLC); Antsiranana, 7 km N of Joffreville, 12°27'S49°14'E, 426 m, dryforest, MT, M. Irwin, R. Harin'Hala leg., MA- 01-07-16, 1♀ (CAS); Antsiranana, Montagne d'Ambre National Park, 12°30'52''S49°10'53''E, 960 m, 19.III–6.IV.2001, MT, R. Harin'Hala leg., MA-01-01A- 09, 1♀ (MOLC); same locality label, 4–19.III.2001, MA-01-01A-08, 1♀ (CAS); same locality label, 21–28.I.2001, MT, M.E. Irwin, E.I. Schlinger & R. Harin'Hala leg., MA-01-01A-01, 1♀ (CAS); Antsiranana, Montagne d'Ambre National Park, 12°31.13'S49°10.45'E, 1125 m, 14–30.V.2001, MT, R. Harin'Hala leg., MA-01-01D-11, 1♀ (CAS); Fianarantsoa, Haute-Matsiatra Region, Andringitra-Ambalavao, Cirque Boby, 2500 m, 13.I.1958, R. Paulian leg., 1♀ (MNHN); Fianarantsoa, Ranomafana National Park, Talatakely, 21°15'S47°25'E, 850 m, 14.IV.1998, 1♀ (CAS); Fianarantsoa, Ranomafana, 15.III.1994, 1♀ (ISNB); 7 km W of Ranomafana, 900 m, 2♀♀ (FSAE, USNM); Fianarantsoa, Ranomafana National Park, Vohiparara, at broken bridge, 21°13.57'S47°22.19'E, 1110 m, 4– 12.II.2002, MT, high altitude rainforest, R. Harin'Hala leg., MA-02-09A-15, 1♀ (CAS); Fianarantsoa, Alakamisy, 21.XI.2006, in forest, C. Canepari leg., 1♀ (MCSNG); Fianarantsoa, Ranomafana National Park, Belle Vue at Ta- latakely, 21°15.99'S47°25.21'E, 1020 m, 26.II–10.III.2003, MT, secondary tropical forest, R. Harin'Hala leg., MA-02-09C-55, 1♀ (MOLC); same locality label, 14–21.I.2002, MA-02-09C-12, 1♀ (CAS); same locality label 22–28.XI.2001, MA-02-09C-04, 1♀ (CAS); Fianarantsoa, Ranomafana National Park, radio tower at forest edge, 21°15.05'S47°24.43'E, 1130 m, 12–19.III.2002, MT, mixed tropical forest, R. Harin'Hala leg., MA-02-09B-20, 1♀ (MOLC); same locality label, 15–21.XII.2001, MA-02-09B-07, 1♀ (CAS); same locality label, 22–28.XI.2001, MA-02-09B-04, 1♀ (MOLC); same locality label, 27.IV–7.V.2003, MA-02-09B-59, 1♀ (CAS); Fianarantsoa, Be- fotaka–Midongy National Park, Papango, 27.7 km S Midongy-South Mt. Papango, 23°50.07'S46°57.49'E, 940 m, 13–19.XI.2006, MT, B.L. Fisher et al . leg., BLF14774, 1♀ (CAS); Fianarantsoa, Agnalazaha Forest Reserve, Ma- habo, 42.9 km 215° SW of Farafangana, 23°11.38'S47°43.23'E, 20 m, 18.IV.2006, MT, littoral rainforest, B.L. Fisher et al. leg., BLF13822, 1♀ (CAS); Fianarantsoa, 50 km S of Farafangana, Mahabo Mananivo, Ampitavanan- ima Forest, 23°07'79''S47°43'02''E, 34 m, 26.I–4.II.2007, MT, in low altitude littoral forest, M. Irwin, F. Parker, R. Harin'Hala leg., MG-35-03, 1♀ (CAS); Fianarantsoa, Miandritsara Forest, 40 km S Ambositra, 20°47.56'S47°10.54'E, 825 m, 25.X–4.XI.2006, MT, low altitude rainforest, M. Irwin, R. Harin'Hala leg., MA-29-56, 1♀ (CAS); Fianarantsoa, near Isalo National Park, at stream east of Interpretive Center, 22°37.60'S45°21.49'E, 750 m, 10–17.III.2002, MT, in open area, MA-02-11A-19, R. Harin'Hala leg., 1♀ (CAS); same locality label, 15–22.IV.2002, MA-02-11A-24, 1♀ (CAS); same locality label, 30.XI–7.XII.2001, MA-02-11A-05, 1♀ (CAS); same locality label, 19–26.I.2002, MA-02-11A-12, 1♀ (CAS); Fianarantsoa, near Isalo National Park, dry wash east of Interpretive Center, 22°37.60'45°21.49'E, 885 m, 9–23.VIII.2002, MT, in open area, MA-02-11B-37, R. Harin'Hala leg., 1♀ (CAS); same locality label, 7–22.IX.2002, MA-02-11B-39, 2♀♀ (CAS); same locality label, 3–17.X.2002, MA-02- 11B-41, 1♀ (CAS); Fianarantsoa, Analalava Forest, 29.6 km 280° W Ranohira, 22°35.30'S45°07.42'E, 700 m, 1– 5.II.2003, tropical dry forest, beating low vegetation, Fisher, Griswold et al . leg., BLF7384, 1♀ (CAS); Mahajanga, Tsingy de Bemaraha National Park, 2.5 km 62° ENE Bekopaka, Ankidrodroa River, 19°07.56'S44°48.53'E, 100 m, 11–15.XI.2001, sifted litter, tropical dry forest on Tsingy, Fisher, Griswold et al . leg., BLF 4340, 1♀ (CAS); Maha- janga, Bale de Baly National Park, 12.4 km 337° NNW Soalala, 16°00.36'S45°15.54'E, 10 m, MT, tropical dry forest, Fisher, Griswold et al . leg., BLF 6811, 2♀♀ (MOLC); Mahajanga, Beanka Forest Reserve, 50.2 km E Main- tirano, 18°01.35'S44°03.02'E, 250 m, 19–26.X.2009, YPT, tropical dry forest on tsingy, B.L. Fisher et al . leg., BLF22788, 3♀♀ (2 in CAS, 1 in MOLC); Mahajanga, Maintirano District, Asondrodava dry forest, 15 km N of Maintirano, 17°57.92'S44°2.13'E, 200 ft., 7–14.XII.2007, MT, dry forest, M. Irwin, R. Harin'Hala leg., MG-41-11, 1♀ (CAS); same locality label, 3–10.XII.2007, MG-43A-09, 1♀ (CAS); same locality label, 8–15.X.2007, 1♀ (CAS); Mahajanga, Analamanitra Forest, 14 km NE Misinjo, 15°55'S45°57'E, 65 ft., 23–30.X.2007, MT, dense dry forest, M. Irwin, R. Harin'Hala leg., MG-38-06, 1♀ (CAS); same locality label, 20.XI–4.XII.2007, MG-38-09, 2♀♀ (CAS); Mahajanga, Ambatofolaka, Namoroka, 53 km from Soalala, 3 km N Vilanandro village, 16°28'04''S45°23'48''E, 400 ft., 21–28.IX.2007, dense dry forest, MT, M. Irwin, R. Harin'Hala leg., MG-40A-01, 1♀ (CAS); Mahajanga, Ampijoroa National Park, 160 km N of Maevatanana on RN 04, 16°19'16''S46°48'80''E, 43 m, 17– 24.VIII.2003, MT, deciduous forest, R. Harin'Hala leg., MA-25-11, 1♀ (CAS); same locality label, 12–19.X.2003, MA-25-19, 1♀ (CAS); same locality label, 28.IX–5.X.2003, MA-25-17, 2♀♀ (CAS); Mahajanga, Namoroka Na- tional Park, 16.9 km 317° NW Vilanandro, 16°24.24'S45°18.36'E, 100 m, 12–16.XI.2002, YPT, tropical dry forest, Fisher, Griswold et al . leg., BLF 6536, 2♀♀ (CAS, MOLC); Mahajanga, Namoroka National Park, 16.40494S45.31294E, 21–31.X.2016, MT, C. Villemant leg., 1♀ (MNHN); Mahajanga, Namoroka National Park, 23– 25.X.2016, YPT, C. Villemant leg., 1♀ (MNHN); Mahajanga, Sofia District, 45 km S Antsohihy, 5 km W Anjiaman- girana, Analagnambe dry degraded forest, 15°09.42'S47°47.05'E, 97 m, 2–6.I.2011, MT, MG-61-04, 1♀ (MOLC); Toamasina, Manakambahiny-E, 17°45'S48°43'E, 1♀ (FSAE); Toamasina, 25 km W of Morarano- Chrome, 17°45'S47°59'E, 1♀ (FSAE); Toamasina, Morarano-Chrome, II.1992, 1♀ (ISNB); Toamasina, Foulpointe, 1♀ (FSAE); Toamasina, Ivondro, 1♀ (MNHN); Toamasina, Rogez (= Andekaleka), VI.1957, Pierre Andria Robinson leg., 1♀ (MNHN); Toamasina, Akirindro Mt., 7.6 km 341° NNW Ambinanitelo, 15°17.18'S49°32.54'E. 600 m, 17– 21.III.2003, YPT, rainforest, Fisher, Griswold et al . leg., BLF8255, 2♀♀ (CAS); Toamasina, Betampona Reserve, Camp Rendrirendry, 34.1 km 332° Toamasina, 17°55.26'S49°11.59'E, 390 m, 28.XI.2005, YPT, rainforest, Brian L. Fisher et al . leg., BLF13124, 1♀ (CAS); Toamasina, Zahamena National Park, Onibe River, 17°45'33''S48°51'17''E, 780 m, 21–23.II.2009, rainforest, YPT, BLF22170, B.L. Fisher et al . leg., 1♀ (CAS); Toamasina, 9.2 km WSW Befingotra, Anjanaharibe-sud Reserve, 14°45'S49°28'E, 1280 m, 13–21.XI.1994, B.L. Fisher leg., 1♀ (DEUCD); Toliara, Bekily, 2♀♀ (MNHN, MRAC); Toliara, Beroboka, 1♀ (NHMUK); Toliara, Beroboka village, 45 km NE Morondava, 19°58.65'S44°39.92'E, 420 ft., 3–11.VIII.2008, MT, Antsarongaza gallery forest, MG-45B- 33, M. Irwin, R. Harin'Hala leg., 1♀ (MOLC); Toliara, Belalanda, 1♀ (FSAE); Toliara, Mahafaly Plateau, 6.2 km 74° ENE of Itampolo, 24°39.13'S43°59.48'E, 80 m, MT, spiny forest thicket, 21–25.II.2002, Fisher, Griswold et al . leg., BLF5759, 1♀ (MOLC); Toliara, Androimpano Forest, 5 km E of Itampolo, 24°39.02'S43°57.79'E, 130 m, 3–16.VII.2009, MT, path in dry forest, M. Irwin, R. Harin'Hala leg., MG-54C-33, 1♀ (CAS); Toliara, Behara, 1♀ (MRAC); Toliara, Fort-Dauphin, 1♀ (MRAC); Toliara, Cap S.te Marie Special Reserve, 74 km S of Tsihombe, 25°35'26''S45°09'78''E, 37 m, 18–26.V.2003, MT, spiny bush, M. Irwin, F. Parker & R. Harin'Hala leg., MA- 02- 23-27, 1♀ (CAS); same locality label, 25.II–7.III.2003, 3♀♀ (2 in CAS, 1 in MOLC); Toliara, Forêt de Kirindy, 15.5 km 64° ENE Marofandilia, 10–17 September 2008, 20°04'09''S044°39'38''E, B.L.Fisher, MT, tropical dry forest, 30 m, BLF18285 _45, 2♀♀ (CAS); Toliara, Mikea Forest, NW of Manombo, 22°54.22'S43°28.53'E, 30 m, 23.IX–3.X.2002, MT, deciduous dry forest, R. Harin'Hala leg., MA-02-18A-34, 1♀ (CAS); same locality label, 6–17.VII.2003, MA-02-18A-61, 1♀ (CAS); Toliara, Andohahela National Park, Tsimelahy, parcelle II, 24°56.21'S46°37.60'E, 180 m, 30.X–9.XI.2003, MT, transitional forest, M. Irwin, F. Parker & R. Harin'Hala leg., MA- 02-20- 44, 1♀ (CAS); same locality label, 5–15.I.2004, MA- 02-20-51, 1♀ (CAS); Toliara, Beza Mahafaly Reserve, par- celle I near research station, 23°41.19'S44°35.46'E, 165 m, 23.III–2.IV.2003, MT, dry deciduous forest, R. Harin'Hala leg., MA-02-14A-54, 1♀ (MOLC); same locality label, 1–8.II.2002, MA-02-14A-14, 1♀ (CAS); same locality label, 8-15.II.2002, MA-02-14A-15, 1♀ (CAS); Toliara, Tsimanampetsotsa National Park, Mitoho Cave, 6.4 km 77° ENE Efoetse, 17.4 km 170° S Beheloka, 24°02.50'S43°45.11'E, 40 m, 18–22.III.2002, MT, spiny forest thicket, Fisher, Griswold et al . leg., BLF 6159, 1♀ (CAS); Toliara, Berenty Special Reserve, 8 km NW Amboasary, 25°00.40'S46°18.20'E, elev 85 m, 17–24.III.2003, M.Irwin, F. Parker & R. Harin'Hala, MT, spiny forest, MA- 02- 22-19, 1♀ (CAS); same locality label, 5–15.II.2003, MA- 02-22-15, 1♀ (CAS); same locality label, 15–25.II.2003, MA- 02-22-16, 1♀ (CAS); same locality label, 16–27.XII.2002, MA- 02-22-10, 1♀ (CAS); Toliara, Berenty, 1♀ (NHMUK); Toliara, Berenty, 12 km NW Amboasary, 5–15.V.1983, J.S. Noyes & M.C. Day leg., 2♀♀ (NHMUK); Toliara, Zombitse National Park, near ANGAP office, 22°53.19'S44°41.53'E, 840 m, 9–19.XI.2001, MT, deciduous spiny forest, M.E. Irwin, F.D. Parker & R. Harin'Hala leg., MA-02-13A-02, 1♀ (CAS); Toliara, Kalambatritra Special Reserve, Ampanihy, 23°27'49''S46°27'47''E, 1270 m, 9–10.II.2009, YPT, montane rainforest, B.L. Fisher et al . leg., BLF21567, 1♀ (CAS); Toliara, Ranobe, 23°02.783'S43°36.617'E, 20 m, 27.X–2.XI.2003, MT, spiny forest thicket, MGF081, Frontier Wilderness Project, 1♀ (CAS); Toliara, Ranobe, 23°02.367'S43°36.617'E, 20 m, 8–15.XI.2003,MT, gallery forest,MGF084, Frontier Wilderness Project, 1♀ (CAS); Toliara, Fiherenana, 23°13.351'S43°52.853'E, 65 m, 5–10.VIII.2003, MT, degraded riparian forest close to water, MGF075, Frontier Wilderness Project, 1♀ (CAS); Toliara, Fiherenana, 23°10.37'S43°57.39'E, 100 m, 31.X–6.XI.2002, MT, gallery forest, MGF045, Frontier Wilderness Project, 4♀♀ (CAS); same locality label, 22–31.X.2002, MGF042, 2♀♀ (CAS, MOLC); Toliara, Manderano, 23°31.39'S44°5.18'E, 70 m, 15–29.V.2002, MT, edge of marsh, 5 m from road, gallery forest, MGF032, Frontier Wilderness Project, 2♀♀ (CAS); Toliara, Makay Mts., 21°13.11'S, 45°19.26'E, 500 m, 24.XI-12.XII.2010, MT, gallery forest, sandy soil, B.L. Fisher et al . leg., BLF25357, 1♀ (CAS); Toliara, Vohi- basia National Park, 44 km NE of Sakaraha, Ambakaka gallery/palm forest, 22°40'S44°50'E, 600 m, 26.XII.2010 - 3.I.2011, M. Irwin, R. Harin'Hala leg., MG-60-12, 1♀ (CAS). SOUTH AFRICA: Limpopo, 13 km W of Beitbridge, Altyddroog Farm, 22°11'S29°53'E, 8.III.1990, C.D. Eardley leg. (SANC); Western Cape, Walker Bay Nature Reserve, 34°27.41'S19°21.39'E, 24.I–21.II.1998, YPT, S. van Noort & B. Fisher leg., 1♀ (SAMC); Western Cape, Gamkaberg Nature Reserve, 33°43.663'S21°57.600'E, 940 m, 10.IX–4.XI.2009, MT, Rooiberg Sandstone Fynbos, S. van Noort leg., 1♀ (SAMC). UGANDA: Uganda, 1♀ (OLL); Kibale National Park, Kanyawara, Makerere Uni- versity Biological Field Station, 0°33.836'N30°21.700'E, 1523 m, 4–26.VIII.2008, YPT, primary mid–altitude rainforest, near stream, S. van Noort leg., 2♀♀ (SAMC). YEMEN: 12 km NW Manakhah, 21.VIII–28.X.2002, MT, N. 7143, A. van Harten leg., 1♀ (MOLC). Hosts . Unknown. Distribution . Botswana, Cameroon, Ethiopia, Madagascar, South Africa, Uganda, Yemen. : Published as part of Olmi, Massimo, Copeland, Robert S. & Noort, Simon Van, 2019, Dryinidae of the Afrotropical region (Hymenoptera, Chrysidoidea), pp. 1-619 in Zootaxa 4630 (1) on pages 342-345, DOI: 10.11646/zootaxa.4630.1.1, http://zenodo.org/record/3336635 : {"references": ["Ceballos, G. (1936) Estudios sobre Anteoninae de Madagascar (Hym., Bethyl.). EOS, 12, 43 - 64.", "Rey del Castillo, C. & Izquierdo, I. (1989) Tipos de especies de Himenopteros descritas por G. Ceballos, en el Museo Nacional de Ciencias Naturales de Madrid. EOS, 65, 251 - 264.", "Benoit, P. L. G. (1954) Monographie des Dryinides Malgaches (Hym. - Acul.). Memoires de l'Institut Scientifique de Madagascar, Serie E, 4, 383 - 430.", "Olmi, M. (1984) A revision of the Dryinidae (Hymenoptera). Memoirs of the American Entomological Institute, 37, I-XII + 1 - 1913.", "Olmi, M. (1994 c) New species of Dryinidae from Madagascar (Hymenoptera Chrysidoidea). Frustula entomologica, New Series, 17, 1 - 12.", "Azevedo, C. O., Madl, M. & Olmi, M. (2010) A Catalogue of the Bethylidae, Chrysididae, Dryinidae, Embolemidae, Sclerogibbidae and Scolebythidae (Hymenoptera: Chrysidoidea) of the Malagasy Subregion. Linzer biologische Beitrage, 42, 845 - 918.", "Olmi, M., van Noort, S. & Guglielmino, A. (2016) Contribution to the knowledge of Afrotropical Dryinidae, Embolemidae and Sclerogibbidae (Hymenoptera), with description of new species from Central African Republic and Uganda. ZooKeys, 578, 45 - 95. https: // doi. org / 10.3897 / zookeys. 578.7820"]}