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feed conversion efficiency feeding Alopex lagopus Scophthalmus maximus animal breeding aquaculture effective population size feed conversion ratio feet fish foxes fur animals genetic trend group dominance dynamics growth heritabiliteetti heritability kalanviljely kalat kotieläinjalostus piikkikampela ruokinta selection selective breeding sinikettu turbot turkiseläimet vesiviljely vulpes lagopus welfare

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