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500 km 800 km 98 %) fidelity Biological Sciences not elsewhere classified Developmental Biology Ecology Environmental Sciences not elsewhere classified Evolutionary Biology annual cycle annual cycles birds stopped breeding ground confront hazards throughout direct migrations along div >< p documented dozens effective management en route </ faster migration hudson bay left winter grounds loop migrations mackenzie delta many long migration routes migration speed migratory pathways multiple years numenius phaeopus </ populations spend