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carbon glacier retreat moss moss bacterial communities moss traits nitrogen nitrogen fixation plant–soil–microbe interactions cyanobacterial abundance and composition primary succession soil soil bacterial communities subarctic ecosystems 500 Naturwissenschaften und Mathematik570 Biowissenschaften; Biologie570 Biowissenschaften; Biologie Behavior and Systematics Biological Soil Crust (BSC) Biological Soil Crust BSC Climate Change Climate change Ecology Ecology, Evolution, Behavior and Systematics Evolution FOS Biological sciences SDG 12 - Responsible Consumption and Production SDG 13 - Climate Action SDG 14 - Life Below Water SDG 6 - Clean Water and Sanitation ddc:570 nitrogen (N) fixation nitrogen N fixation