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cogradient variation countergradient variation latitudinal gradient Rana arvalis [SDE.BE]Environmental Sciences/Biodiversity and Ecology local adaptation colonization Q ST – F ST comparison amphibians climate change daptation ecological genetics life history evolution life-history traits Ecology, Evolution, Behavior and Systematics Life sciences QST-FST comparison envir medicine and health care Genetics anthro-se life history traits Adaptive divergence Amphibians Colonization Divergent selection Evolutionsbiologi Genetic drift Genome Report Lagopus muta

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