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altitudinal gradient mycorrhizal fungi Arctic biodiversity Arters utbredning Betulaceae Dicotyledons Ecology, Evolution, Behavior and Systematics Root-associated fungi Species distribution angiosperms arctic region areas and regions areas created by nature arktiska regionen biologi biologisk mångfald biology birches björkar björkväxter broad-leaved trees change downy birch ecology ekologi entiteter environmental changes events events and action