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brood size costs-benefits defending behaviour disease transmission prey spectrum urbanization GPS tracking arctic waterfowl cultivated grassland migratory fuelling carry-over effects recruitment 60801 Animal Behaviour Ecology Environmental Science FOS Biological sciences Branta bernicla bernicla Terschelling The Netherlands Wadden Sea 5.29 E 53.38 N Life Science Lolium perenne Lolium Brood size Costs-benefits Defending behaviour Disease transmission Pasture

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