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boreal dissolved organic carbon Arctic particulate organic carbon permafrost carbon wildfire permafrost Ecology [SDE.BE]Environmental Sciences/Biodiversity and Ecology carbon cycling coastal erosion coastal erosion Supplementary material for this article is available cryosphere and high-latitude processes nutrients and nutrient cycling rivers thermokarst 1172 Environmental sciences CLIMATE-CHANGE Ekologi Global change Lakes Phytoplankton Spatial Temporal carbon nutrients ARCTIC TUNDRA BOREAL FOREST CO2 Climate Research