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Aquatic Science ecosystem dynamics environmental drivers food web mass-balance modeling primary production variability sequential depletion trophic flows Ecology Environmental Sciences not elsewhere classified Evolutionary Biology FOS Biological sciences FOS Chemical sciences Inorganic Chemistry Plant Biology Geology [SDE.BE]Environmental Sciences/Biodiversity and Ecology [SDU.STU.OC]Sciences of the Universe [physics]/Earth Sciences/Oceanography benthos bottom trawling environment marine biology seafloor svalbard Animal Science and Zoology Arctic Article Barents Sea Ecology, Evolution, Behavior and Systematics General Agricultural and Biological Sciences