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Canis lupus Holocene Y-chromosome wolf Carnivora Leptospira borgpetersenii Hybridization Iberian Peninsula Nuclear markers dog nuclear markers spirochete Articles Aullidos Bayesian hierarchical-site-occupancy models Continental patterns Ecology Gestión de lobos Health, Toxicology and Mutagenesis Homesites Howling Human population density Human-dominated landscapes Human-made structures Landscape-level decision mak-ing Large carnivores Life sciences Marcaje territorial Modelos bayesianos jerárquicos de ocupación de sitio Monitoring optimization