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Hymenogastraceae Agaricales Agaricomycetes Basidiomycota Fungi Hebeloma Arctic distribution High Arctic Low Arctic mycorrhizal hosts new species pruned median joining networks A.H. Smith Arctic-alpine ITS ectomycorrhizal fungal biodiversity systematics Agaricomycetidae Agaricomycotina Astraeus hygrometricus Gigaspermataceae Meteora Palavascia Synchytriales Unikonta Hebeloma leucosarx Hebeloma alpinum Hebeloma aurantioumbrinum Hebeloma hiemale

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