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5 cm bls 65 cm Biological Sciences not elsewhere classified Ecology Environmental Sciences not elsewhere classified Genetics Microbiology Science Policy active layer active layer belonged ammonia oxidation arctic consists assembled genomes showed bacterial taxa biodiversity decreased significantly carbon soil comparative metagenomics component systems contrasting depths derived polysaccharide degradation functionally different gene abundance genes encoding genetic potential improve prediction main electron acceptor metagenomes revealed enrichment microbial activity microbial communities prevalent microbes identified